Uruguaytherium
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Uruguaytherium
''Uruguaytherium'' is an extinct genus of astrapotherid mammal from the Late Oligocene to Early Miocene of South America. It was named by the Argentinean paleontologist Lucas Kraglievich in 1928, from a fragmentary fossil found in the Fray Bentos Formation of the department of Río Negro in Uruguay, and the type species is ''U. beaulieui''.''Uruguaytherium''
at .org
The related genera '''' and ''

Uruguaytheriinae
Astrapotheriidae is an extinct family of herbivorous South American land mammals that lived from the Late Eocene (Mustersan SALMA) to the Middle Miocene (Laventan SALMA) . The most derived of the astrapotherians, they were also the largest and most specialized mammals in the Tertiary of South America. There are two sister taxa: Eoastrapostylopidae and Trigonostylopidae. Around 1900, Argentine paleontologist Florentino Ameghino described eight Colhuehuapian (Early Miocene) species from specimens he found south of Lake Colhué Huapi in Patagonia and grouped them into three genera: '' Parastrapotherium'', ''Astrapotherium'', and '' Astrapothericulus''. It was obvious to Ameghino that these species represented a great diversity, ranging in size from a peccary to a rhinoceros, but his description was based entirely on fragmentary and not always comparable dental remains. Other expeditions to Patagonia have subsequently recovered considerably more complete materials. Genera According ...
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Astrapotheriidae
Astrapotheriidae is an extinct family of herbivorous South American land mammals that lived from the Late Eocene (Mustersan SALMA) to the Middle Miocene (Laventan SALMA) . The most derived of the astrapotherians, they were also the largest and most specialized mammals in the Tertiary of South America. There are two sister taxa: Eoastrapostylopidae and Trigonostylopidae. Around 1900, Argentine paleontologist Florentino Ameghino described eight Colhuehuapian (Early Miocene) species from specimens he found south of Lake Colhué Huapi in Patagonia and grouped them into three genera: '' Parastrapotherium'', ''Astrapotherium'', and '' Astrapothericulus''. It was obvious to Ameghino that these species represented a great diversity, ranging in size from a peccary to a rhinoceros, but his description was based entirely on fragmentary and not always comparable dental remains. Other expeditions to Patagonia have subsequently recovered considerably more complete materials. Genera According ...
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Deseadan
The Deseadan ( es, Deseadense) age is a period of geologic time (29.0–21.0 Ma) within the Oligocene epoch of the Paleogene to the Early Miocene epoch of the Neogene, used more specifically within the SALMA classification of South America. It follows the Tinguirirican and precedes the Colhuehuapian age. Etymology The age is named after the Deseado Formation of the Deseado Massif in eastern Patagonia, Argentina. Formations Fossils Correlations The Deseadan South American land mammal age (SALMA) is equivalent to the Arikareean in the North American land mammal age (NALMA) and the Harrisonian in the 2000 version of the classification. It overlaps with the Hsandagolian The Hsandagolian age is a period of geologic time (33.9 – 23.03 Ma) within the Oligocene epoch of the Paleogene used more specifically with Asian Land Mammal Ages. It follows the Kekeamuan and precedes the Tabenbulakian age. The Ulangochuian ... of Asia and the MP 25 zone of Europe, the Waitaki ...
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Xenastrapotherium
''Xenastrapotherium'' is an extinct genus of astrapothere, a type of hoofed herbivorous mammal, native to South America, which lived in the Middle to Late Miocene period, typically during the Laventan stage. It is a member of the family Astrapotheriidae in the subfamily Uruguaytheriinae, large astrapotheres, equipped with a trunk-like nose and protruding teeth, similar to the elephants, but their tusks were the canine teeth, not the incisors. ''Xenastrapotherium'' (named after the Greek word ''xenos'' "strange" add to the genus ''Astrapotherium'', "lightning beast") was a genus widely distributed in northern South America, in contrast to other species of astrapotheres which lived in the area of the Southern Cone of the continent. It differed from other astrapotheres by having two lower incisors on each side of the jaw and the tusks have a pronounced longitudinal curvature, although their general shape and size are probably very similar to ''Astrapotherium'', whose weight would be ...
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Trigonostylops
''Trigonostylops'' is an extinct genus of South American meridiungulatan ungulate, from the Late Paleocene to Late Eocene (Itaboraian to Tinguirirican in the SALMA classification) of South America (Argentina and Peru) and Antarctica (Seymour Island). It is the only member of the family Trigonostylopidae. Description A complete skull of the type species, ''T. wortmani'', has been found, and it has been classified as an astrapothere based on its large lower incisors. Phylogeny Cladogram based in the phylogenetic analysis published by Vallejo Pareja ''et al''., 2015, showing the position of ''Trigonostylops'':Vallejo Pareja et al., 2015 Distribution Fossils of ''Trigonostylops'' have been found in:''Trigonostylops''
at

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Granastrapotherium
''Granastrapotherium'' is an extinct genus of ungulate mammals, described from remains found in rocks of the Honda Group in the Tatacoa Desert, in the Colombian departments of Huila and Tolima, at the Miocene fossil site La Venta. The only species formally recognized is ''Granastrapotherium snorki'' (from Spanish, ''gran'', "great"; ''Astrapotherium'', "lightning beast"; and snorkel, breathing tube, in reference to the trunk). Remains found in Bolivia and Peru, seem to belong to ''Granastrapotherium'' or a very similar animal. Description This astrapothere differs from its coeval, the uruguaytheriine astrapotheriid ''Xenastrapotherium'' by their larger size, between 3 and 4 tonnes, with tusks about one meter long, making it one of the largest representatives of Astrapotheria, only surpassed by some species of ''Parastrapotherium''. Other differences include the presence of only one premolar, the lack of incisors in both jaws and the disposition of the canine tooth, which are ...
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Astraponotus
''Astraponotus'' is an extinct genus of astrapotheriids. It lived during the Middle-Late Eocene (in the Mustersan and Tinguirirican of the South American land mammal ages (SALMA), 48-33.9 million years ago) and its fossil remains have been found in the Sarmiento Formation of Argentina, South America.''Astraponotus''
at .org


Description

Unlike most astrapotheres, ''Astraponotus'' was equipped with an unusually high, short, narrow skull. The nasal bones were quite withdrawn, which suggests the presence of a short proboscis. Other unusual features of ''Astraponotus'' includes the extreme reduction of the

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Cladogram
A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to descendants, nor does it show how much they have changed, so many differing evolutionary trees can be consistent with the same cladogram. A cladogram uses lines that branch off in different directions ending at a clade, a group of organisms with a last common ancestor. There are many shapes of cladograms but they all have lines that branch off from other lines. The lines can be traced back to where they branch off. These branching off points represent a hypothetical ancestor (not an actual entity) which can be inferred to exhibit the traits shared among the terminal taxa above it. This hypothetical ancestor might then provide clues about the order of evolution of various features, adaptation, and other evolutionary narratives about ance ...
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Phylogenetic Analysis
In biology, phylogenetics (; from Greek φυλή/ φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups of organisms. These relationships are determined by Computational phylogenetics, phylogenetic inference methods that focus on observed heritable traits, such as DNA sequences, protein amino acid sequences, or morphology. The result of such an analysis is a phylogenetic tree—a diagram containing a hypothesis of relationships that reflects the evolutionary history of a group of organisms. The tips of a phylogenetic tree can be living taxa or fossils, and represent the "end" or the present time in an evolutionary lineage. A phylogenetic diagram can be rooted or unrooted. A rooted tree diagram indicates the hypothetical common ancestor of the tree. An unrooted tree diagram (a network) makes no assumption about the ancestral line, and does n ...
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Eoastrapostylops
''Eoastrapostylops'' is an extinct genus of astrapothere that lived during the Late Paleocene in what is now Argentina. Description This animal was small in size; the skull was 9 centimeters long and the total length of the animal probably just exceeded half a meter. ''Eoastrapostylops'' possessed a short muzzle ; its canines were well developed but not yet derived into large fangs, as was the case in later South American ungulates such as ''Trigonostylops'', and the nasal bones were not retracted, indicating the absence of the typical proboscis exhibited by later forms. The dentition of ''Eoastrapostylops'' recalls that of its later relative, ''Trigonostylops''. molars and premolars were low-crowned and lophoselenodont-shaped. The fourth premolar was molarized, while both the fourth upper premolar and the third upper molar were triangularly shaped and lacked an hypoconus. Classification ''Eoastrapostylops riolorense'' was first described in 1981, based on fossils found in the ...
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Albertogaudrya
''Albertogaudrya'' is an extinct genus of astrapotherian mammal that lived in present-day Salta, Argentina (, paleocoordinates ) during the Eocene (Casamayoran SALMA The South American land mammal ages (SALMA) establish a geologic timescale for prehistoric South American fauna beginning 64.5 Ma during the Paleocene and continuing through to the Late Pleistocene (0.011 Ma). These periods are referred to as a ...) .''Albertogaudrya''
at Fossilworks.org
Fossils of ''Albertogaudrya'' have been found in the Lumbrera and
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Tetragonostylops
''Tetragonostylops'' is an extinct genus of mammal, related to Astrapotheria. It lived during the Late Paleocene, and its fossils were discovered in South America. Description This genus is only known from incomplete remains, notably a well-preserved mandible and a fragmentary skull. Like other astrapotheres, ''Tetragonostylops'' may have had a strong elongated body, and a long, flat skull. The skull was 20 centimeters long, and the entire body was supposedly one meter long. Its mandible was strong, and endowed with an elongated cylindrical symphysis. The mandibular condyle was elevated, and the coronoid process was high, but lower than in its relative ''Trigonostylops''. The shape of their molars was different, with ''Tetragonostylops'' having more squared molars (hence his name, ''Tetragon-'' meaning "four corners"), and had an infraorbital canal, which does not appear in ''Trigonostylops''. The molars were brachyodont, like those of ''Trigonostylops'', and the canines were st ...
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