True Toad
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True Toad
A true toad is any member of the family Bufonidae, in the order Anura (frogs and toads). This is the only family of anurans in which all members are known as toads, although some may be called frogs (such as harlequin frogs). The bufonids now comprise more than 35 genera, ''Bufo'' being the best known. History Bufonidae is thought to have originated in South America. Some studies date the origin of the group to after the breakup of Gondwana, about 78 - 98 million years ago in the Late Cretaceous. In contrast, other studies have dated the origin of the group to the early Paleocene. The bufonids likely radiated out of South America during the Eocene, with the entire radiation occurring during the Eocene to Oligocene, marking an extremely rapid divergence likely facilitated by the Paleogene's changing climatic conditions. Taxonomy The following phylogeny of most genera in the family is based on Portik and Papenfuss, 2015:, Chan ''et al.'', 2016, Chandramouli ''et al.'', 2016 ...
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Paleocene
The Paleocene, ( ) or Palaeocene, is a geological epoch (geology), epoch that lasted from about 66 to 56 mya (unit), million years ago (mya). It is the first epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), Era. The name is a combination of the Ancient Greek ''palaiós'' meaning "old" and the Eocene Epoch (which succeeds the Paleocene), translating to "the old part of the Eocene". The epoch is bracketed by two major events in Earth's history. The K–Pg extinction event, brought on by Chicxulub impact, an asteroid impact and possibly volcanism, marked the beginning of the Paleocene and killed off 75% of living species, most famously the non-avian dinosaurs. The end of the epoch was marked by the Paleocene–Eocene Thermal Maximum (PETM), which was a major climatic event wherein about 2,500–4,500 gigatons of carbon were released into the atmosphere and ocean systems, causing a spike in global temperatures and ocean acidification. In the Pal ...
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Oreophrynella
''Oreophrynella'', commonly known as bush toads, is a genus of true toads native to the tepuis of southern Venezuela and adjacent Guyana. The distribution of some species is restricted to a couple of tepuis or even a single tepui, as in the case of '' Oreophrynella weiassipuensis'', which occurs on Wei-Assipu-tepui. Description Species of the genus ''Oreophrynella'' are small frogs, less than in snout–vent length. They are characterized by opposable digits of the foot, dorsal skin that bears tubercules, and direct development (i.e., there is no free-living larval stage). The presence of opposable digits, unique among bufonids, in combination with an extension of the interdigital integument and the relative length/orientation of the digits, is likely to be an adaptation to facilitate life on rocky tepui summits and an exaptation to arboreality The genus also displays cranial Standard anatomical terms of location are used to unambiguously describe the anatomy of animals, i ...
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Sclerophrys
''Sclerophrys'' is a genus of "true toads", family Bufonidae, native to Africa and the southern Arabian Peninsula. Originally, all of these species were classified in the genus ''Bufo''. The genus, originally named ''Amietophrynus'', was split due to large enough taxonomic divergence. Ohler and Dubois showed in 2016 that ''Sclerophrys capensis'' Tschudi, 1838 is the same species as '' Bufo regularis rangeri'' Hewitt, 1935, the type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen ... of ''Amietophrynus''. Because the former name is older, the implication is that ''Amietophrynus'' is a junior synonym of ''Sclerophrys''. Species The following species are recognized in the genus ''Sclerophrys''. References Amphibian genera Amphibians of Africa Amphibians of Asia Ta ...
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Poyntonophrynus
''Poyntonophrynus'', also known as pygmy toads, are a genus consisting of ten true toad species native to Sub-Saharan Africa. Originally, all ''Poyntonophrynus'' species were included in the genus ''Bufo''. The genus was split due to large enough taxonomic divergence as evidenced by molecular markers. The genus is named in honour of , South African herpetologist, with ''phrynus'' being Greek for toad. Taxonomy and systematics ''Poyntonophrynus'' corresponds to the former '' Bufo vertebralis'' group. ''Poyntonophrynus'' is the sister taxon of ''Mertensophryne''. Some molecular studies have suggested polyphyly of ''Poyntonophrynus'', but the conclusion is likely to have been caused by sequences representing a different species than assumed. Description ''Poyntonophrynus'' are small toads. They lack a tarsal fold, and their parotoid glands are indistinct and flattened. The tympanum is small but distinct. Species The species in this genus are: * '' Poyntonophrynus beiranus'' (Lover ...
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Didynamipus
The four-digit toad or dwarf toad (''Didynamipus sjostedti'') is a species of toad in the family Bufonidae. It is found in Bioko (Equatorial Guinea), southwestern Cameroon, and southeastern Nigeria. It is the only species in the genus ''Didynamipus''. Etymology The specific name ''sjostedti'' honours Bror Yngve Sjöstedt, a Swedish entomologist and ornithologist who collected the type series. Description ''Didynamipus sjostedti'' are very small toads: males grow to and females to in snout–vent length. The snout is sharp in males but rounded in females; males also have a ridge on the top the snout that gives it an upturned appearance. The tympanum is absent. The hands and feet are reduced, with the two outer fingers and toes and the innermost toes present only as tubercles. No webbing is present. The dorsum is brown with lighter and darker mottling. Habitat and conservation ''Didynamipus sjostedti'' occur on forest edges and in clearings in moist forest at elevations of ...
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Incilius
''Incilius'' is genus of toads in the true toad family, Bufonidae. They are sometimes known as the Central American toads or Middle American toads and are found in southern USA, Mexico, Central America, and northern Pacific South America (Colombia and Ecuador). They are an ecologically and biogeographically diverse group of toads, including micro-endemic species such as '' Incilius spiculatus'' that are restricted to undisturbed cloud forests, and widespread lowland species such as ''Incilius valliceps'' that predominantly occur in disturbed habitats. Taxonomy and systematics This genus was first described in 1863 by Edward Drinker Cope who designated the type species as ''Incilius coniferus''. This proved unpopular and these toads were known under the genus ''Bufo'' until the early 2000s. The current delineation of the genus follows Mendelson ''et al''. (2011) who brought ''Cranopsis''/''Cranophryne''/''Ollotis'' and ''Crepidius''/''Crepidophryne'' into synonymy with ''Inciliu ...
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Anaxyrus
''Anaxyrus'' is a genus of true toads in the family Bufonidae. The genus is endemic to North and Central America. Some authors consider ''Anaxyrus'' to be a subgenus within ''Bufo ''Bufo'' is a genus of true toads in the amphibian family Bufonidae. As traditionally defined, it was a wastebasket genus containing a large number of toads from much of the world, but following taxonomic reviews most of these have been moved to ...''. Species References External links * * * Further reading * Tschudi JJ (1845). "''Reptilium conspectus quae in Republica Peruana reperiuntur et pleraque observata vel collecta sunt in itinere''". ''Archiv für Naturgeschichte'' 11 (1): 150–170. (''Anaxyrus'', new genus, p. 170). (in Latin) {{Taxonbar, from=Q2279985 Taxa named by Johann Jakob von Tschudi ...
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Rhinella
''Rhinella'', commonly known as South American toads, beaked toads or Rio Viejo toads, is a genus of true toads native to Neotropical parts of Mexico, Central and South America. Additionally, the cane toad has been introduced to Australia, the Caribbean, the Philippines and elsewhere. Originally, all species of the genus ''Rhinella'' were included in the genus ''Bufo'', then they were split into the genera ''Chaunus'' and ''Rhamphophryne''. However, ''Chaunus'' and ''Rhamphophryne'' are now considered synonyms of ''Rhinella''. Etymology * ''Rhinella'' means ‘little nose’, from ''rhino-'' (), the combining form of the Ancient Greek ' (, ‘nose’) and the Latin diminutive A diminutive is a root word that has been modified to convey a slighter degree of its root meaning, either to convey the smallness of the object or quality named, or to convey a sense of intimacy or endearment. A (abbreviated ) is a word-formati ... suffix '' -ella''. * ''Chaunus'' is the Latinised form ...
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Rhaebo
''Rhaebo'' is a genus of true toads, family Bufonidae, from Central and South America. They are distributed from Honduras to northern South America including the Amazonian lowlands. Common name Cope toads has been suggested for them. Taxonomy The genus was removed from the synonymy of ''Bufo'' in 2006; an alternative view has been to treat it as a subgenus of ''Bufo''. At present, it is widely recognized as a genus. ''Andinophryne'', consisting of three species, was recognized as a separate genus until 2015 when it was found out that its recognition rendered ''Rhaebo'' paraphyletic. An alternative to synonymizing it with ''Rhaebo'' would have been to erect a new genus for '' Rhaebo nasicus'', but this would have caused difficulty in assigning species without molecular data to correct genus. Description ''Rhaebo'' are characterized as lacking cephalic crests, having omosternum, distinctively wide sphenethmoid, prominent and notched exoccipital condyles, and yellowish-orange sk ...
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Peltophryne
''Peltophryne'' is a genus of true toads in the family Bufonidae, from the Greater Antilles (Cuba, Isla de Juventud, Hispaniola, Puerto Rico). With ten endemic species, Cuba hosts the highest diversity. Hispaniola has three endemics and Puerto Rico and the Virgin Islands combined has one. Description ''Peltophryne'' range is size from the relatively small '' Peltophryne cataulaciceps'' with a snout–vent length (SVL) of to the large '' Peltophryne peltocephala'' with SVL of . The skull is as long as wide and contains some unique osteological features (thickened dermal tissue covering the snout and usually ossified into a pair of rostral bones, and squamosal-maxillary articulation). These are considered to be derived characters that set these toads apart from other bufonids. Taxonomy The genus was erected by Leopold Fitzinger in 1843, but placed in synonymy with ''Bufo'' by Albert Günther in 1859. Subsequent work has considered ''Peltophryne'' either as a valid genus, a subgen ...
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Nannophryne
''Nannophryne'' is a small genus of true toads, family Bufonidae, from South America. They are found in central Andean Peru south to southern Chile and adjacent Argentina. Taxonomy The genus was placed in synonymy with ''Bufo'' by George Albert Boulenger in 1894, but it was resurrected in 2006 by Frost when splitting the then very large ''Bufo'' into smaller, monophyletic genera. This change was controversial but has now largely been accepted. The alternative, treating ''Nannophryne'' as a subgenus In biology, a subgenus (plural: subgenera) is a taxonomic rank directly below genus. In the International Code of Zoological Nomenclature, a subgeneric name can be used independently or included in a species name, in parentheses, placed between t ... of ''Bufo'', still has some following. Species There are four species in this genus: * '' Nannophryne apolobambica'' (De la Riva, Ríos, and Aparicio, 2005) * '' Nannophryne cophotis'' (Boulenger, 1900) * '' Nannophryne corynetes'' (Due ...
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Dendrophryniscus
''Dendrophryniscus'' is a genus of true toads in the family Bufonidae, sometimes known as tree toads. They are endemic to the Atlantic Forest of Brazil. Taxonomy Until 2012, the genus ''Dendrophryniscus'' included the species now separated to the genus '' Amazophrynella''. This was based on molecular genetic evidence that indicated deep divergence between an Amazonian and an Atlantic Forest clade, the latter retaining the name ''Dendrophryniscus'' whereas the former was described as a new genus ''Amazonella'', later amended to ''Amazophrynella'' because of homonymy. Description The species of ''Dendrophryniscus'' are small to medium-sized toads measuring in snout–vent length. Their body form resembles those of the genus ''Atelopus''. The hind limbs are well developed. The parotoid glands are absent, as are external tympana. The skin is uniformly granulose to warty. Dorsal coloration is cryptic. With the exception of '' Dendrophryniscus leucomystax'', species of the genus ''D ...
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