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''Oreophrynella'', commonly known as bush toads, is a genus of
true toads A true toad is any member of the family Bufonidae, in the order Anura (frogs and toads). This is the only family of anurans in which all members are known as toads, although some may be called frogs (such as harlequin frogs). The bufonids now ...
native to the
tepui A tepui , or tepuy (), is a table-top mountain or mesa found in South America, especially in Venezuela and western Guyana. The word tepui means "house of the gods" in the native tongue of the Pemon, the indigenous people who inhabit the Gran ...
s of southern Venezuela and adjacent Guyana. The distribution of some species is restricted to a couple of tepuis or even a single tepui, as in the case of '' Oreophrynella weiassipuensis'', which occurs on Wei-Assipu-tepui.


Description

Species of the genus ''Oreophrynella'' are small frogs, less than in snout–vent length. They are characterized by opposable digits of the foot, dorsal skin that bears tubercules, and direct development (i.e., there is no free-living larval stage). The presence of opposable digits, unique among bufonids, in combination with an extension of the interdigital integument and the relative length/orientation of the digits, is likely to be an adaptation to facilitate life on rocky
tepui A tepui , or tepuy (), is a table-top mountain or mesa found in South America, especially in Venezuela and western Guyana. The word tepui means "house of the gods" in the native tongue of the Pemon, the indigenous people who inhabit the Gran ...
summits and an
exaptation Exaptation and the related term co-option describe a shift in the function of a trait during evolution. For example, a trait can evolve because it served one particular function, but subsequently it may come to serve another. Exaptations are commo ...
to
arboreal Arboreal locomotion is the locomotion of animals in trees. In habitats in which trees are present, animals have evolved to move in them. Some animals may scale trees only occasionally, but others are exclusively arboreal. The habitats pose num ...
ity The genus also displays cranial simplification in the form of cranial fontanelles and absence of the quadratojugal, which may be driven by a reduction of developmental costs, increase in flexibility, and reduction of body weight. The cranial simplification combined with the shortening of the vertebral column and shift towards a partly
firmisternal Firmisternal denotes a morphology of the pectoral girdle of frogs. In firmisternal girdles, the epicoracoid cartilages are fused anteriorly and posteriorly, and usually have an omosternum present. This is in contrast to the other predominant morpho ...
girdle may be adaptations to the peculiar tumbling behaviour displayed by ''Oreophrynella''.


Species

The following species are recognised in the genus ''Oreophrynella'':


References


External links

* taxo
''Oreophrynella''
at http://www.eol.org. * Taxo
''Oreophrynella''
at https://www.itis.gov/index.html. (Accessed: May 1, 2008). * Taxo
''Oreophrynella''
at http://data.gbif.org/welcome.htm {{Taxonbar, from=Q134726 Amphibian genera Amphibians of South America Taxa named by George Albert Boulenger Amphibians of the Tepuis