Trogloxene
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Trogloxene
Trogloxenes or subtroglophiles, also called cave guests, are animal species which periodically live in underground habitats such as caves or at the very entrance, but cannot live exclusively in such habitats."Cave Plants and Animals". 9 December Among many scientists, trogloxenes and subtroglophile have slightly different but closely related meanings, with the former covering species that are occasional visitors to underground habitat and the latter species that live more permanently there, but have to go outside (for example, to find food). Both these are in contrast to troglobites, which strictly live in underground habitats. Examples of trogloxene/subtroglophile species are bats, rats, raccoons and some opiliones (this last group also has fully troglobitic species). Several extinct trogloxenes are known like cave bears, cave lions, cave leopards, and cave hyenas. Indications trusted by geologists and archaeologists combine to show that these animals lived there in the latter pa ...
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Stygofauna
Stygofauna are any fauna that live in groundwater systems or aquifers, such as caves, fissures and vugs. Stygofauna and troglofauna are the two types of subterranean fauna (based on life-history). Both are associated with subterranean environments – stygofauna are associated with water, and troglofauna with caves and spaces above the water table. Stygofauna can live within freshwater aquifers and within the pore spaces of limestone, calcrete or laterite, whilst larger animals can be found in cave waters and wells. Stygofaunal animals, like troglofauna, are divided into three groups based on their life history - stygophiles, stygoxenes, and stygobites. # Stygophiles inhabit both surface and subterranean aquatic environments, but are not necessarily restricted to either. # Stygoxenes are like stygophiles, except they are defined as accidental or occasional presence in subterranean waters. Stygophiles and stygoxenes may live for part of their lives in caves, but don't complete th ...
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Species
In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. Other ways of defining species include their karyotype, DNA sequence, morphology, behaviour or ecological niche. In addition, paleontologists use the concept of the chronospecies since fossil reproduction cannot be examined. The most recent rigorous estimate for the total number of species of eukaryotes is between 8 and 8.7 million. However, only about 14% of these had been described by 2011. All species (except viruses) are given a two-part name, a "binomial". The first part of a binomial is the genus to which the species belongs. The second part is called the specific name or the specific epithet (in botanical nomenclature, also sometimes i ...
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Troglobite
A troglobite (or, formally, troglobiont) is an animal species, or population of a species, strictly bound to underground habitats, such as caves. These are separate from species that mainly live in above-ground habitats but are also able to live underground (eutroglophiles), and species that are only cave visitors ( subtroglophiles and trogloxenes). Land-dwelling troglobites may be referred to as troglofauna, while aquatic species may be called stygofauna, although for these animals the term ''stygobite'' is preferable. Troglobites typically have evolutionary adaptations to cave life. Examples of such adaptations include slow metabolism, reduced energy consumption, better food usage efficiency, decrease or loss of eyesight (anophthalmia), and depigmentation (absence of pigment in the integument). Conversely, as opposed to lost or reduced functions, many species have evolved elongated antenna and locomotory appendages, in order to better move around and respond to environmental s ...
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Raccoon
The raccoon ( or , ''Procyon lotor''), sometimes called the common raccoon to distinguish it from other species, is a mammal native to North America. It is the largest of the procyonid family, having a body length of , and a body weight of . Its grayish coat mostly consists of dense underfur, which insulates it against cold weather. Three of the raccoon's most distinctive features are its extremely dexterous front paws, its facial mask, and its ringed tail, which are themes in the mythologies of the indigenous peoples of the Americas relating to the animal. The raccoon is noted for its intelligence, as studies show that it is able to remember the solution to tasks for at least three years. It is usually nocturnal and omnivorous, eating about 40% invertebrates, 33% plants, and 27% vertebrates. The original habitats of the raccoon are deciduous and mixed forests, but due to their adaptability, they have extended their range to mountainous areas, coastal marshes, and urban ...
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Opiliones
The Opiliones (formerly Phalangida) are an order of arachnids colloquially known as harvestmen, harvesters, harvest spiders, or daddy longlegs. , over 6,650 species of harvestmen have been discovered worldwide, although the total number of extant species may exceed 10,000. The order Opiliones includes five suborders: Cyphophthalmi, Eupnoi, Dyspnoi, Laniatores, and Tetrophthalmi, which were named in 2014. Representatives of each extant suborder can be found on all continents except Antarctica. Well-preserved fossils have been found in the 400-million-year-old Rhynie cherts of Scotland, and 305-million-year-old rocks in France. These fossils look surprisingly modern, indicating that their basic body shape developed very early on, and, at least in some taxa, has changed little since that time. Their phylogenetic position within the Arachnida is disputed; their closest relatives may be the mites (Acari) or the Novogenuata (the Scorpiones, Pseudoscorpiones, and Solifugae). Althou ...
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Cave Bear
The cave bear (''Ursus spelaeus'') is a prehistoric species of bear that lived in Europe and Asia during the Pleistocene and became extinct about 24,000 years ago during the Last Glacial Maximum. Both the word "cave" and the scientific name ''spelaeus'' are used because fossils of this species were mostly found in caves. This reflects the views of experts that cave bears may have spent more time in caves than the brown bear, which uses caves only for hibernation. Taxonomy Cave bear skeletons were first described in 1774 by Johann Friedrich Esper, in his book ''Newly Discovered Zoolites of Unknown Four Footed Animals''. While scientists at the time considered that the skeletons could belong to apes, canids, felids, or even dragons or unicorns, Esper postulated that they actually belonged to polar bears. Twenty years later, Johann Christian Rosenmüller, an anatomist at Leipzig University, gave the species its binomial name. The bones were so numerous that most researcher ...
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Lion
The lion (''Panthera leo'') is a large Felidae, cat of the genus ''Panthera'' native to Africa and India. It has a muscular, broad-chested body; short, rounded head; round ears; and a hairy tuft at the end of its tail. It is sexually dimorphic; adult male lions are larger than females and have a prominent mane. It is a social species, forming groups called ''prides''. A lion's pride consists of a few adult males, related females, and cubs. Groups of female lions usually hunt together, preying mostly on large ungulates. The lion is an apex predator, apex and keystone predator; although some lions scavenge when opportunities occur and have been known to hunt Human, humans, lions typically don't actively seek out and prey on humans. The lion inhabits grasslands, savannas and shrublands. It is usually more diurnality, diurnal than other wild cats, but when persecuted, it adapts to being active nocturnality, at night and crepuscular, at twilight. During the Neolithic period, the li ...
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European Ice Age Leopard
''Panthera pardus spelaea'', sometimes called the European Ice Age leopard or Late Pleistocene leopard, is a fossil leopard subspecies, which roamed Europe in the Late Pleistocene. The youngest known bone fragments date to about 32,000 to 26,000 years ago, and are similar in size to modern leopard bones. Taxonomy Several fossil bones from the Early, Middle and Late Pleistocene were described and proposed as different leopard subspecies: *''P. p. antiqua'' *''P. p. begoueni'' *''P. p. sickenbergi'' *''P. p. vraonensis'' These are now considered junior synonyms of ''P. p. spelaea''. Description The European Ice Age leopard's skull was medium-long, and its characteristics are closest to the ''Panthera pardus tulliana'' subspecies. An apparent depiction of this leopard in the Chauvet Cave shows a coat pattern similar to that of modern leopards but with a unspotted belly, presumably white. Like other mammals, leopards from the cold glacial periods of the Late Pleistocene are usua ...
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Cave Hyena
The cave hyena (''Crocuta crocuta spelaea''), also known as the Ice Age spotted hyena, was a paleosubspecies of spotted hyena which ranged from the Iberian Peninsula to eastern Siberia. It is one of the best known mammals of the Ice Age and is well represented in many European bone caves. The cave hyena was a highly specialised animal, with its progressive and regressive features being more developed than in its modern African relative. It preyed on large mammals (primarily wild horses, steppe bison and woolly rhinoceros), and was responsible for the accumulation of hundreds of large Pleistocene mammal bones in areas including horizontal caves, sinkholes, mud pits and muddy areas along rivers. The cause of the cave hyena's extinction is not fully understood, though it could have been due to a combination of factors, including climate change and competition with other predators. Description and paleoecology The main difference between the spotted hyena and the cave hyena lies in ...
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Interglacial
An interglacial period (or alternatively interglacial, interglaciation) is a geological interval of warmer global average temperature lasting thousands of years that separates consecutive glacial periods within an ice age. The current Holocene interglacial began at the end of the Pleistocene, about 11,700 years ago. Pleistocene During the 2.5 million years of the Pleistocene, numerous glacials, or significant advances of continental ice sheets, in North America and Europe, occurred at intervals of approximately 40,000 to 100,000 years. The long glacial periods were separated by more temperate and shorter interglacials. During interglacials, such as the present one, the climate warms and the tundra recedes polewards following the ice sheets. Forests return to areas that once supported tundra vegetation. Interglacials are identified on land or in shallow epicontinental seas by their paleontology. Floral and faunal remains of species pointing to temperate climate and indicating a s ...
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Last Glacial Maximum
The Last Glacial Maximum (LGM), also referred to as the Late Glacial Maximum, was the most recent time during the Last Glacial Period that ice sheets were at their greatest extent. Ice sheets covered much of Northern North America, Northern Europe, and Asia and profoundly affected Earth's climate by causing drought, desertification, and a large drop in sea levels. Based on changes in position of ice sheet margins dated via terrestrial cosmogenic nuclides and radiocarbon dating, growth of ice sheets commenced 33,000 years ago and maximum coverage was between 26,500 years and 19–20,000 years ago, when deglaciation commenced in the Northern Hemisphere, causing an abrupt rise in sea level. Decline of the West Antarctica ice sheet occurred between 14,000 and 15,000 years ago, consistent with evidence for another abrupt rise in the sea level about 14,500 years ago. Glacier fluctuations around the Strait of Magellan suggest the peak in glacial surface area was constrained to betwee ...
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Upper Paleolithic
The Upper Paleolithic (or Upper Palaeolithic) is the third and last subdivision of the Paleolithic or Old Stone Age. Very broadly, it dates to between 50,000 and 12,000 years ago (the beginning of the Holocene), according to some theories coinciding with the appearance of behavioral modernity in early modern humans, until the advent of the Neolithic Revolution and agriculture. Anatomically modern humans (i.e. ''Homo sapiens'') are believed to have emerged in Africa around 300,000 years ago, it has been argued by some that their ways of life changed relatively little from that of archaic humans of the Middle Paleolithic, until about 50,000 years ago, when there was a marked increase in the diversity of Artefact (archaeology), artefacts found associated with modern human remains. This period coincides with the most common date assigned to early human migrations, expansion of modern humans from Africa throughout Asia and Eurasia, which contributed to the Neanderthal extinction, ex ...
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