Thelxiope
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Thelxiope
''Thelxiope'' is a genus of Cambrian and Ordovician arthropod. Four named species are known, the type species ''T. palaeothalassia'' is known from the Burgess Shale, Canada ''T. holmani'' is from the Wheeler Shale of Utah, ''Thelxiope tangi'' from the Linyi Lagerstätte of Shandong, China, and ''T. spinosa'', which is known from both the Linyi Lagerstätte and the Wheeler Shale. An indeterminate species is also known from the Ordovician (Floian) Fezouata Formation in Morocco. It is a member of Mollisoniida, alongside close relatives ''Mollisonia, Corcorania'' and ''Urokodia.'' They are suggested to be stem-chelicerates The subphylum Chelicerata (from New Latin, , ) constitutes one of the major subdivisions of the phylum Arthropoda. It contains the sea spiders, horseshoe crabs, and arachnids (including harvestmen, scorpions, spiders, solifuges, ticks, and mite .... References Burgess Shale fossils Prehistoric arthropod genera {{paleo-arthropod-stub Cambrian genu ...
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Fezouata Formation
The Fezouata Formation or Fezouata Shale is a geological formation in Morocco which dates to the Early Ordovician.Fezouata Formation
at Fossilworks.org
Fezouata Shale
at Fossilworks.org

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Urokodia
''Urokodia aequalis'' is an extinct genus of arthropod from the early Cambrian. The taxon is only known from the Maotianshan Shales of China based on some 15 specimens. Its segmentation resembles that of a millipede and it possessed head and tail shields with thorny spikes. It has some similarities to the arthropod ''Mollisonia'' that is known from both the Burgess Shale of Canada and the Kaili biota of China. Recently, the taxon has been considered a member of the order Mollisoniida, alongside ''Mollisonia, Thelxiope,'' and ''Corcorania,'' the group are suggested to be stem-Chelicerata, chelicerates. References Urokodia aequalis Arthropod from Maotianshan Shales
- URL retrieved September 11, 2010 Prehistoric crustaceans Cambrian arthropods Maotianshan shales fossils {{paleo-arthropod-stub Cambrian genus extinctions ...
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Mollisonia
''Mollisonia'' is an extinct genus of Cambrian arthropod. Species are known from the Burgess Shale, Langston Formation, and Wheeler Shale of North America, as well as the Chengjiang Biota of China. Twenty-one specimens of ''Mollisonia'' are known from the Greater Phyllopod bed, where they comprise less than 0.1% of the community. Remains possibly attributable to the genus are also known from the Ordovician Fezouata Formation of Morocco and Bøggild Fjord Formation Greenland. An observation published in 2019 suggests this genus is a basal chelicerate, closer to crown group Chelicerata than members of Habeliida. It is suggested to be closely related to '' Corcorania'', ''Urokodia'', and ''Thelxiope'', which together form the clade Mollisoniida, which are thought to be closely related to Chelicerata The subphylum Chelicerata (from New Latin, , ) constitutes one of the major subdivisions of the phylum Arthropoda. It contains the sea spiders, horseshoe crabs, and arachnids (in ...
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Chelicerata
The subphylum Chelicerata (from New Latin, , ) constitutes one of the major subdivisions of the phylum Arthropoda. It contains the sea spiders, horseshoe crabs, and arachnids (including Opiliones, harvestmen, scorpions, spiders, Solifugae, solifuges, Parasitiformes, ticks, and Acariformes, mites, among many others), as well as a number of extinct lineages, such as the eurypterids (sea scorpions) and chasmataspidids. The Chelicerata originated as marine animals in the Middle Cambrian period; the first confirmed chelicerate fossils, belonging to ''Sanctacaris'', date from Burgess Shale, 508 million years ago. The surviving marine species include the four species of xiphosurans (horseshoe crabs), and possibly the 1,300 species of Pycnogonida, pycnogonids (sea spiders), if the latter are indeed chelicerates. On the other hand, there are over 77,000 well-identified species of air-breathing chelicerates, and there may be about 500,000 unidentified species. Like all arth ...
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Burgess Shale Fossils
The fossils of the Burgess Shale, like the Burgess Shale itself, formed around 505 million years ago in the Mid Cambrian period. They were discovered in Canada in 1886, and Charles Doolittle Walcott collected over 65,000 specimens in a series of field trips up to the alpine site from 1909 to 1924. After a period of neglect from the 1930s to the early 1960s, new excavations and re-examinations of Walcott's collection continue to reveal new species, and statistical analysis suggests that additional discoveries will continue for the foreseeable future. Stephen Jay Gould's book '' Wonderful Life'' describes the history of discovery up to the early 1980s, although his analysis of the implications for evolution has been contested. The fossil beds are in a series of shale layers, averaging and totalling about in thickness. These layers were deposited against the face of a high undersea limestone cliff. All these features were later raised up above current sea level during the cr ...
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Cambrian
The Cambrian Period ( ; sometimes symbolized C with bar, Ꞓ) was the first geological period of the Paleozoic Era, and of the Phanerozoic Eon. The Cambrian lasted 53.4 million years from the end of the preceding Ediacaran Period 538.8 million years ago (mya) to the beginning of the Ordovician Period mya. Its subdivisions, and its base, are somewhat in flux. The period was established as "Cambrian series" by Adam Sedgwick, who named it after Cambria, the Latin name for 'Cymru' (Wales), where Britain's Cambrian rocks are best exposed. Sedgwick identified the layer as part of his task, along with Roderick Murchison, to subdivide the large "Transition Series", although the two geologists disagreed for a while on the appropriate categorization. The Cambrian is unique in its unusually high proportion of sedimentary deposits, sites of exceptional preservation where "soft" parts of organisms are preserved as well as their more resistant shells. As a result, our understanding of the Ca ...
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Ordovician
The Ordovician ( ) is a geologic period and System (geology), system, the second of six periods of the Paleozoic Era (geology), Era. The Ordovician spans 41.6 million years from the end of the Cambrian Period million years ago (Mya) to the start of the Silurian Period Mya. The Ordovician, named after the Celtic Britons, Welsh tribe of the Ordovices, was defined by Charles Lapworth in 1879 to resolve a dispute between followers of Adam Sedgwick and Roderick Murchison, who were placing the same Rock (geology), rock beds in North Wales in the Cambrian and Silurian systems, respectively. Lapworth recognized that the fossil fauna in the disputed Stratum, strata were different from those of either the Cambrian or the Silurian systems, and placed them in a system of their own. The Ordovician received international approval in 1960 (forty years after Lapworth's death), when it was adopted as an official period of the Paleozoic Era by the International Union of Geological Sciences, Intern ...
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Arthropod
Arthropods (, (gen. ποδός)) are invertebrate animals with an exoskeleton, a Segmentation (biology), segmented body, and paired jointed appendages. Arthropods form the phylum Arthropoda. They are distinguished by their jointed limbs and Arthropod cuticle, cuticle made of chitin, often Mineralization (biology), mineralised with calcium carbonate. The arthropod body plan consists of segments, each with a pair of appendages. Arthropods are bilaterally symmetrical and their body possesses an exoskeleton, external skeleton. In order to keep growing, they must go through stages of moulting, a process by which they shed their exoskeleton to reveal a new one. Some species have wings. They are an extremely diverse group, with up to 10 million species. The haemocoel, an arthropod's internal cavity, through which its haemolymph – analogue of blood – circulates, accommodates its interior Organ (anatomy), organs; it has an open circulatory system. Like their exteriors, the internal or ...
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Burgess Shale
The Burgess Shale is a fossil-bearing deposit exposed in the Canadian Rockies of British Columbia, Canada. It is famous for the exceptional preservation of the soft parts of its fossils. At old (middle Cambrian), it is one of the earliest fossil beds containing soft-part imprints. The rock unit is a black shale and crops out at a number of localities near the town of Field in Yoho National Park Yoho National Park ( ) is a National Parks of Canada, national park of Canada. It is located within the Canadian Rockies, Rocky Mountains along the western slope of the Continental Divide of the Americas in southeastern British Columbia, bordered ... and the Kicking Horse Pass. Another outcrop is in Kootenay National Park 42 km to the south. History and significance The Burgess Shale was discovered by palaeontologist Charles Doolittle Walcott, Charles Walcott on 30 August 1909, towards the end of the season's fieldwork. He returned in 1910 with his sons, daughter, and wif ...
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Wheeler Shale
The Wheeler Shale (named by Charles Walcott) is a Cambrian ( 507  Ma) fossil locality world-famous for prolific agnostid and ''Elrathia kingii'' trilobite remains (even though many areas are barren of fossils) and represents a Konzentrat-Lagerstätte. Varied soft bodied organisms are locally preserved, a fauna (including ''Naraoia'', ''Wiwaxia'' and ''Hallucigenia'') and preservation style (carbonaceous film) normally associated with the more famous Burgess Shale. As such, the Wheeler Shale also represents a Konservat-Lagerstätten. Together with the Marjum Formation and lower Weeks Formation, the Wheeler Shale forms of limestone and shale exposed in one of the thickest, most fossiliferous and best exposed sequences of Middle Cambrian rocks in North America. At the type locality of Wheeler Amphitheater, House Range, Millard County, western Utah, the Wheeler Shale consists of a heterogeneous succession of highly calcareous shale, shaley limestone, mudstone and thi ...
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Linyi Lagerstätte
Linyi () is a prefecture-level city in the south of Shandong province, China. As of 2011, Linyi is the largest prefecture-level city in Shandong, both by area and population, Linyi borders Rizhao to the east, Weifang to the northeast, Zibo to the north, Tai'an to the northwest, Jining to the west, Zaozhuang to the southwest, and the province of Jiangsu to the south. The name of the city Linyi () literally means "close to the Yi River". The city recently expanded along the Yi River to Nanfang, now called the Beicheng New Area, under the slogan "Grand Linyi, Beautiful Linyi, New Linyi". Multiple recreational parks were built, along with new school campuses etc. The development is a consequence of a series of governmental projects, including relocate the city government, which is expected to stimulate the economy. The population was 11,018,365 at the 2020 census, of which 3,651,868 lived in the built-up area made up of Lanshan District, Luozhuang District and Hedong District, a ...
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Shandong
Shandong ( , ; ; alternately romanized as Shantung) is a coastal province of the People's Republic of China and is part of the East China region. Shandong has played a major role in Chinese history since the beginning of Chinese civilization along the lower reaches of the Yellow River. It has served as a pivotal cultural and religious center for Taoism, Chinese Buddhism and Confucianism. Shandong's Mount Tai is the most revered mountain of Taoism and a site with one of the longest histories of continuous religious worship in the world. The Buddhist temples in the mountains to the south of the provincial capital of Jinan were once among the foremost Buddhist sites in China. The city of Qufu is the birthplace of Confucius and was later established as the center of Confucianism. Confucianism developed from what was later called the Hundred Schools of Thought from the teachings of the Chinese philosopher Confucius. Shandong's location at the intersection of ancient and modern n ...
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