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Tyto Gigantea
''Tyto gigantea'' is an extinct barn owl from what is now Gargano Gargano (, Gargano Apulian Italo-Romance arˈgæːnə is a historical and geographical sub-region in the province of Foggia, Apulia, southeast Italy, consisting of a wide isolated mountain massif made of highland and several peaks and forming ..., Italy, dating back to the late Miocene. From its remains, ''T. gigantea'' is suggested to have been as large as or larger than the Eurasian eagle-owl (''Bubo bubo''). This species seems to have lived at the same time as the closely related owl, ''Tyto robusta''. That would represent a single lineage of owls adapting to the largest size of their prey on the island, such as the great Erinaceidae, erinaceomorph of Gargano, ''Deinogalerix'', and rodents.Mlíkovský, J. (1998)A new barn owl (Aves: Strigidae) from the early Miocene of Germany, with comments on the fossil history of the Tytoninae ''Journal für Ornithologie'', 139(3), 247-261. References

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Miocene
The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recent" because it has 18% fewer modern marine invertebrates than the Pliocene has. The Miocene is preceded by the Oligocene and is followed by the Pliocene. As Earth went from the Oligocene through the Miocene and into the Pliocene, the climate slowly cooled towards a series of ice ages. The Miocene boundaries are not marked by a single distinct global event but consist rather of regionally defined boundaries between the warmer Oligocene and the cooler Pliocene Epoch. During the Early Miocene, the Arabian Peninsula collided with Eurasia, severing the connection between the Mediterranean and Indian Ocean, and allowing a faunal interchange to occur between Eurasia and Africa, including the dispersal of proboscideans into Eurasia. During the ...
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Tyto
''Tyto'' is a genus of birds consisting of true barn owls, grass owls and masked owls that collectively make up all the species within the subfamily Tytoninae of the barn owl family, Tytonidae. Taxonomy The genus ''Tyto'' was introduced in 1828 by the Swedish naturalist Gustaf Johan Billberg with the western barn owl as the type species. The name is from the Ancient Greek ''tutō'' meaning "owl". The barn owl (''Tyto alba'') was formerly considered to have a global distribution with around 28 subspecies. In the list of birds maintained by Frank Gill, Pamela Rasmussen and David Donsker on behalf of the International Ornithological Committee (IOC) the barn owl is now split into four species: the western barn owl (''Tyto alba'') (10 subspecies), the American barn owl (''Tyto furcata'') (12 subspecies), the eastern barn owl (''Tyto javanica'') (7 subspecies) and the Andaman masked owl (''Tyto deroepstorffi''). This arrangement is followed here. Some support for this split was provide ...
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Barn Owl
The barn owl (''Tyto alba'') is the most widely distributed species of owl in the world and one of the most widespread of all species of birds, being found almost everywhere except for the polar and desert regions, Asia north of the Himalayas, most of Indonesia, and some Pacific Islands. It is also known as the common barn owl, to distinguish it from the other species in its family, Tytonidae, which forms one of the two main lineages of living owls, the other being the typical owls (''Strigidae''). There are at least three major lineages of barn owl: the western barn owl of Europe, western Asia, and Africa; the eastern barn owl of southeastern Asia and Australasia; and the American barn owl of the Americas. Some taxonomic authorities classify barn owls differently, recognising up to five separate species; and further research needs to be done to resolve the disparate taxonomies. There is considerable variation of size and colour among the approximately 28 subspecies, but most ...
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Gargano
Gargano (, Gargano Apulian Italo-Romance arˈgæːnə is a historical and geographical sub-region in the province of Foggia, Apulia, southeast Italy, consisting of a wide isolated mountain massif made of highland and several peaks and forming the backbone of the Gargano Promontory projecting into the Adriatic Sea, the "spur" on the Italian "boot". The high point is Monte Calvo at . Most of the upland area, about , is part of the Gargano National Park, founded in 1991. In this region since 1978 a feud has been fought between the clans of the Società foggiana. The Gargano peninsula is partly covered by the remains of an ancient forest, ''Foresta Umbra'', the only remaining part in Italy of the ancient oak and beech forest that once covered much of Central Europe as well as the Apennine deciduous montane forests ecoregion. The Latin poet Horace spoke of the oaks of ''Garganus'' in ''Ode'' II, ix. Tourism The coast of Gargano houses numerous beaches and tourist facilities, inclu ...
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Eurasian Eagle-owl
The Eurasian eagle-owl (''Bubo bubo'') is a species of eagle-owl that resides in much of Palearctic, Eurasia. It is also called the Uhu and it is occasionally abbreviated to just the eagle-owl in Europe. It is one of the largest species of owl, and females can grow to a total length of 75 cm (30 in), with a wingspan of 188 cm (6 ft 2 in), with males being slightly smaller.''Owls of the World: A Photographic Guide'' by Mikkola, H. Firefly Books (2012), This bird has distinctive ear tufts, with upper parts that are mottled with darker blackish colouring and tawny. The wings and tail are barred. The underparts are a variably hued buff, streaked with darker colouring. The facial disc is not very defined and the orange eyes are distinctive.Penteriani, V., & del Mar Delgado, M. (2019). ''The eagle owl''. Bloomsbury Publishing. Eurasian eagle-owls are found in many habitats, but are mostly birds of Mountain ecosystems, mountainous regions or other Rock (geolog ...
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Tyto Gigantea - Tibiotarsus
''Tyto'' is a genus of birds consisting of true barn owls, grass owls and masked owls that collectively make up all the species within the subfamily Tytoninae of the barn owl family, Tytonidae. Taxonomy The genus ''Tyto'' was introduced in 1828 by the Swedish naturalist Gustaf Johan Billberg with the western barn owl as the type species. The name is from the Ancient Greek ''tutō'' meaning "owl". The barn owl (''Tyto alba'') was formerly considered to have a global distribution with around 28 subspecies. In the list of birds maintained by Frank Gill, Pamela Rasmussen and David Donsker on behalf of the International Ornithological Committee (IOC) the barn owl is now split into four species: the western barn owl (''Tyto alba'') (10 subspecies), the American barn owl (''Tyto furcata'') (12 subspecies), the eastern barn owl (''Tyto javanica'') (7 subspecies) and the Andaman masked owl (''Tyto deroepstorffi''). This arrangement is followed here. Some support for this split was provide ...
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Tyto Robusta
''Tyto robusta'' was a prehistoric barn-owl. It lived at what is now Monte Gargano in Italy, and was an island throughout much of the Neogene when sea levels were higher. The owl's remains date back to the Miocene-Pliocene boundary 5.5 to 5 million years ago. The fossil bones are about 60% as long again as a modern barn owl, giving a total length of about 50–65 cm for ''T. robusta''. This owl provides an interesting case study of evolution and insular gigantism. Evolution, taxonomy and systematics It was found sympatrically with the gigantic relative, '' Tyto gigantea'', which at some 220% of the size of the barn owl grew to some 70–85 cm in length—larger than a Eurasian eagle-owl and maybe up to twice as heavy. Like other apex predators they can be presumed to have occurred at low population densities. Being apparently endemic to Gargano Island and perhaps some other islands in today's Apennines, where one or two huge eagles and at least at times a normal-sized b ...
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Erinaceidae
Erinaceidae is a family in the order Eulipotyphla, consisting of the hedgehogs and moonrats. Until recently, it was assigned to the order Erinaceomorpha, which has been subsumed with the paraphyletic Soricomorpha into Eulipotyphla. Eulipotyphla has been shown to be monophyletic; Soricomorpha is paraphyletic because Soricidae shared a more recent common ancestor with Erinaceidae than with other soricomorphs. Erinaceidae contains the well-known hedgehogs (subfamily Erinaceinae) of Eurasia and Africa and the gymnures or moonrats (subfamily Galericinae) of South-east Asia. This family was once considered part of the order Insectivora, but that polyphyletic order is now considered defunct. Characteristics Erinaceids are generally shrew-like in form, with long snouts and short tails. They are, however, much larger than shrews, ranging from in body length and in weight, in the case of the short-tailed gymnure, up to and in the moonrat. All but one species have five toes in eac ...
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Deinogalerix
''Deinogalerix'' (from Ancient Greek, "terrible/terror" + ''Galerix'') is an extinct genus of gymnure which lived in Italy in the Late Miocene, 7-10 million years ago. The genus was apparently endemic to what was then the island of Gargano, which is now a peninsula. The first specimens of ''Deinogalerix'' were first described in 1972. The genus is in the hedgehog subfamily of gymnures or moon-rats, which are not rats at all, but rather hairy, superficially rat-like relatives of the hedgehog lacking quills. ''Deinogalerix'' had a long, thin, conical face, small pointed ears, a lengthy, tapering tail and long hairs. ''Deinogalerix koenigswaldi''s skull was long and the entire body measured . It occupied the same ecological niche as dogs and cats today, except that it had more predators itself - such as the enormous barn owl '' Tyto gigantea''. It is believed that the species of ''Deinogalerix'' were insectivores, mostly feeding off invertebrates like beetles, dragonflies and ...
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Miocene Birds
The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recent" because it has 18% fewer modern marine invertebrates than the Pliocene has. The Miocene is preceded by the Oligocene and is followed by the Pliocene. As Earth went from the Oligocene through the Miocene and into the Pliocene, the climate slowly cooled towards a series of ice ages. The Miocene boundaries are not marked by a single distinct global event but consist rather of regionally defined boundaries between the warmer Oligocene and the cooler Pliocene Epoch. During the Early Miocene, the Arabian Peninsula collided with Eurasia, severing the connection between the Mediterranean and Indian Ocean, and allowing a faunal interchange to occur between Eurasia and Africa, including the dispersal of proboscideans into Eurasia. During the late ...
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Fossils Of Italy
A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved in amber, hair, petrified wood and DNA remnants. The totality of fossils is known as the ''fossil record''. Paleontology is the study of fossils: their age, method of formation, and evolutionary significance. Specimens are usually considered to be fossils if they are over 10,000 years old. The oldest fossils are around 3.48 billion years old to 4.1 billion years old. Early edition, published online before print. The observation in the 19th century that certain fossils were associated with certain rock strata led to the recognition of a geological timescale and the relative ages of different fossils. The development of radiometric dating techniques in the early 20th century allowed scientists to quantitatively measure the absolute ...
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