Gargano (, Gargano Apulian Italo-Romance arˈgæːnə is a historical and geographical sub-region in the
province of Foggia
The Province of Foggia ( it, Provincia di Foggia ; Foggiano: ) is a province in the Apulia (Puglia) region of southern Italy.
This province is also known as Daunia, after the Daunians, an Iapygian pre-Roman tribe living in Tavoliere plain, and ...
Italy
Italy ( it, Italia ), officially the Italian Republic, ) or the Republic of Italy, is a country in Southern Europe. It is located in the middle of the Mediterranean Sea, and its territory largely coincides with the homonymous geographical ...
, consisting of a wide isolated mountain massif made of highland and several peaks and forming the backbone of the Gargano Promontory projecting into the
Adriatic Sea
The Adriatic Sea () is a body of water separating the Italian Peninsula from the Balkans, Balkan Peninsula. The Adriatic is the northernmost arm of the Mediterranean Sea, extending from the Strait of Otranto (where it connects to the Ionian Sea) ...
, the "spur" on the Italian "boot". The high point is Monte Calvo at . Most of the upland area, about , is part of the Gargano National Park, founded in 1991. In this region since 1978 a feud has been fought between the clans of the
Società foggiana
The Società foggiana, also known as ''Mafia Foggiana'' (''Foggian mafia'') and the fifth mafia (in addition to Cosa Nostra in Sicily, the ’Ndrangheta in Calabria, the Camorra in Campania and the Sacra Corona Unita in greater Apulia, from wh ...
.
The Gargano peninsula is partly covered by the remains of an ancient forest, ''Foresta Umbra'', the only remaining part in Italy of the ancient oak and
beech
Beech (''Fagus'') is a genus of deciduous trees in the family Fagaceae, native to temperate Europe, Asia, and North America. Recent classifications recognize 10 to 13 species in two distinct subgenera, ''Engleriana'' and ''Fagus''. The ''Engl ...
ecoregion
An ecoregion (ecological region) or ecozone (ecological zone) is an ecologically and geographically defined area that is smaller than a bioregion, which in turn is smaller than a biogeographic realm. Ecoregions cover relatively large areas o ...
. The Latin poet
Horace
Quintus Horatius Flaccus (; 8 December 65 – 27 November 8 BC), known in the English-speaking world as Horace (), was the leading Roman lyric poet during the time of Augustus (also known as Octavian). The rhetorician Quintilian regarded his ...
spoke of the oaks of ''Garganus'' in '' Ode'' II, ix.
Tourism
The coast of Gargano houses numerous beaches and tourist facilities, including resorts such as Vieste, Peschici and Mattinata. The two major
salt lake
A salt lake or saline lake is a landlocked body of water that has a concentration of salts (typically sodium chloride) and other dissolved minerals significantly higher than most lakes (often defined as at least three grams of salt per litre). ...
s of Lesina and Varano are located in the northern part of the peninsula. Gargano is the site of the oldest shrine in
Western Europe
Western Europe is the western region of Europe. The region's countries and territories vary depending on context.
The concept of "the West" appeared in Europe in juxtaposition to "the East" and originally applied to the ancient Mediterranean ...
dedicated to the
archangel Michael
Michael (; he, מִיכָאֵל, lit=Who is like El od, translit=Mīḵāʾēl; el, Μιχαήλ, translit=Mikhaḗl; la, Michahel; ar, ميخائيل ، مِيكَالَ ، ميكائيل, translit=Mīkāʾīl, Mīkāl, Mīkhāʾīl), also ...
Triassic
The Triassic ( ) is a geologic period and system which spans 50.6 million years from the end of the Permian Period 251.902 million years ago (Mya), to the beginning of the Jurassic Period 201.36 Mya. The Triassic is the first and shortest period ...
Period, known as "Black Stones" in Lesina, as well as the Sanctuary of San Nazario.
Good Friday
Good Friday is a Christian holiday commemorating the crucifixion of Jesus and his death at Calvary. It is observed during Holy Week as part of the Paschal Triduum. It is also known as Holy Friday, Great Friday, Great and Holy Friday (also Ho ...
Saint Valentine
Saint Valentine ( it, San Valentino; la, Valentinus) was a 3rd-century Roman saint, commemorated in Western Christianity on February 14 and in Eastern Orthodoxy on July 6. From the High Middle Ages, his Saints' Day has been associated with a ...
trail running
Trail running is a sport-activity which combines running, and, where there are steep gradients, hiking, that is run "on any unpaved surface". It is similar to both mountain and fell running (also known as hill running). Mountain running may, ...
ultra distance running
An ultramarathon, also called ultra distance or ultra running, is any footrace longer than the traditional marathon length of . Various distances are raced competitively, from the shortest common ultramarathon of to over . 50k and 100k are both ...
half marathon
A half marathon is a road running event of —half the distance of a marathon. It is common for a half marathon event to be held concurrently with a marathon or a 5K race, using almost the same course with a late start, an early finish or shortcu ...
Late Miocene
The Late Miocene (also known as Upper Miocene) is a sub-epoch of the Miocene Epoch made up of two stages. The Tortonian and Messinian stages comprise the Late Miocene sub-epoch, which lasted from 11.63 Ma (million years ago) to 5.333 Ma.
The ...
to
Early Pliocene
Early may refer to:
History
* The beginning or oldest part of a defined historical period, as opposed to middle or late periods, e.g.:
** Early Christianity
** Early modern Europe
Places in the United States
* Early, Iowa
* Early, Texas
* Early ...
, a highly
endemic
Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found els ...
vertebrate
Vertebrates () comprise all animal taxon, taxa within the subphylum Vertebrata () (chordates with vertebral column, backbones), including all mammals, birds, reptiles, amphibians, and fish. Vertebrates represent the overwhelming majority of the ...
fauna
Fauna is all of the animal life present in a particular region or time. The corresponding term for plants is ''flora'', and for fungi, it is '' funga''. Flora, fauna, funga and other forms of life are collectively referred to as ''biota''. Zoo ...
evolved on what was then Gargano Island due to higher
sea level
Mean sea level (MSL, often shortened to sea level) is an average surface level of one or more among Earth's coastal bodies of water from which heights such as elevation may be measured. The global MSL is a type of vertical datuma standardise ...
s than today. Several of these animals were subject to
island gigantism
Island gigantism, or insular gigantism, is a biological phenomenon in which the size of an animal species isolated on an island increases dramatically in comparison to its mainland relatives. Island gigantism is one aspect of the more general Fos ...
.
The
fossil
A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...
s are found in partially infilled paleokarst fissures across Monte Gargano. The Gargano Island fauna is known as '' Mikrotia'' fauna after an endemic
rodent
Rodents (from Latin , 'to gnaw') are mammals of the Order (biology), order Rodentia (), which are characterized by a single pair of continuously growing incisors in each of the upper and lower jaws. About 40% of all mammal species are roden ...
genus
Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial n ...
of the area. Initially named ''Microtia'', this had to be corrected, because the genus name '' Microtia'' was already used for
butterflies
Butterflies are insects in the macrolepidopteran clade Rhopalocera from the order Lepidoptera, which also includes moths. Adult butterflies have large, often brightly coloured wings, and conspicuous, fluttering flight. The group comprises t ...
.
The surface features of the ancient karst developed in
Mesozoic
The Mesozoic Era ( ), also called the Age of Reptiles, the Age of Conifers, and colloquially as the Age of the Dinosaurs is the second-to-last era of Earth's geological history, lasting from about , comprising the Triassic, Jurassic and Creta ...
limestone
Limestone ( calcium carbonate ) is a type of carbonate sedimentary rock which is the main source of the material lime. It is composed mostly of the minerals calcite and aragonite, which are different crystal forms of . Limestone forms wh ...
. In these,
sediment
Sediment is a naturally occurring material that is broken down by processes of weathering and erosion, and is subsequently transported by the action of wind, water, or ice or by the force of gravity acting on the particles. For example, sa ...
accumulated together with the remains of the local fauna, forming thick layers of reddish, massive or crudely stratified
silt
Silt is granular material of a size between sand and clay and composed mostly of broken grains of quartz. Silt may occur as a soil (often mixed with sand or clay) or as sediment mixed in suspension with water. Silt usually has a floury feel wh ...
y-
sand
Sand is a granular material composed of finely divided mineral particles. Sand has various compositions but is defined by its grain size. Sand grains are smaller than gravel and coarser than silt. Sand can also refer to a soil texture, textur ...
y
clay
Clay is a type of fine-grained natural soil material containing clay minerals (hydrous aluminium phyllosilicates, e.g. kaolin, Al2 Si2 O5( OH)4).
Clays develop plasticity when wet, due to a molecular film of water surrounding the clay part ...
s, known as ''terrae rossae'' ("red
soil
Soil, also commonly referred to as earth or dirt, is a mixture of organic matter, minerals, gases, liquids, and organisms that together support life. Some scientific definitions distinguish ''dirt'' from ''soil'' by restricting the former ...
s"). Through the mid-
Pliocene
The Pliocene ( ; also Pleiocene) is the epoch in the geologic time scale that extends from 5.333 million to 2.58tectonic movement of the Apulian Plate. Others were overlaid by other sediments of
terrestrial
Terrestrial refers to things related to land or the planet Earth.
Terrestrial may also refer to:
* Terrestrial animal, an animal that lives on land opposed to living in water, or sometimes an animal that lives on or near the ground, as opposed to ...
or
freshwater
Fresh water or freshwater is any naturally occurring liquid or frozen water containing low concentrations of dissolved salts and other total dissolved solids. Although the term specifically excludes seawater and brackish water, it does in ...
origin. In this way a buried, partially reworked paleokarst originated.
Later, as the ice ages cycle got underway, sea levels sank and the former island was continentalized. In the cool and
semiarid
A semi-arid climate, semi-desert climate, or steppe climate is a dry climate sub-type. It is located on regions that receive precipitation below potential evapotranspiration, but not as low as a desert climate. There are different kinds of semi-ar ...
conditions of the Early Pleistocene (some 1.8–0.8 mya) a second karstic cycle occurred, producing the neokarst which removed part of the paleokarst fill.
Deinogalerix
''Deinogalerix'' (from Ancient Greek, "terrible/terror" + ''Galerix'') is an extinct genus of gymnure which lived in Italy in the Late Miocene, 7-10 million years ago. The genus was apparently endemic to what was then the island of Gargano, whic ...
'' - 5 species of gymnures ("hairy hedgehogs"), among them the giant ''D. koenigswaldi'' with a skull of c.20 cm length. (Freudenthal, 1972; Butler, 1980)
* ''
Hoplitomeryx
''Hoplitomeryx'' is a genus of extinct deer-like ruminants which lived on the former Gargano Island during the Miocene and the Early Pliocene, now a peninsula on the east coast of South Italy. ''Hoplitomeryx'', also known as "prongdeer", had fi ...
'' - some 5 species of "prongdeer" with five horns and sabre-like upper
canine teeth
In mammalian oral anatomy, the canine teeth, also called cuspids, dog teeth, or (in the context of the upper jaw) fangs, eye teeth, vampire teeth, or vampire fangs, are the relatively long, pointed teeth. They can appear more flattened however ...
. They ranged from tiny to the size of a
red deer
The red deer (''Cervus elaphus'') is one of the largest deer species. A male red deer is called a stag or hart, and a female is called a hind. The red deer inhabits most of Europe, the Caucasus Mountains region, Anatolia, Iran, and parts of wes ...
, and large and small ones apparently occurred at the same time rather than one evolving from the other. (Leinders 1984, van der Geer 2005, van der Geer 2008)
* ''Mikrotia'' - 3 or more species of
murine
The Old World rats and mice, part of the subfamily Murinae in the family Muridae, comprise at least 519 species. Members of this subfamily are called murines. In terms of species richness, this subfamily is larger than all mammal families ex ...
rodent. The largest species, ''M. magna'', had a skull 10 cm long. (Freudenthal, 1976, Parra ''et al.'', 1999)
* ''Paralutra garganensis'' - an endemic species of otter. (Willemsen, 1983)
* '' Prolagus imperialis'' and ''P. apricenicus'' - huge endemic pika species- ''P. imperialis'' was larger than any other known ''Prolagus''.(Mazza, 1987)
* ''Stertomys'' - 5 species of
dormouse
A dormouse is a rodent of the family Gliridae (this family is also variously called Myoxidae or Muscardinidae by different taxonomists). Dormice are nocturnal animals found in Africa, Asia, and Europe. They are named for their long, dormant hib ...
, among them the giant ''S. laticrestatus'' (Daams and Freudenthal, 1985) and four smaller species (Freudenthal and Martín-Suárez, 2006)
* ''Hattomys'' - 3 species of giant hamsters, among them the giant ''H. gargantua''. (Freudenthal, 1985)
Non-endemic mammals found on the island included:
* '' Apodemus gorafensis'' - a field mouse
* A prehistoric species of '' Cricetus''
hamster
Hamsters are rodents (order Rodentia) belonging to the subfamily Cricetinae, which contains 19 species classified in seven genera.Fox, Sue. 2006. ''Hamsters''. T.F.H. Publications Inc. They have become established as popular small pets. The ...
(Freudenthal, 1985)
* ''Megacricetodon'' - another hamster (Freudenthal, 1985)
Bird species occurring at Gargano included (studied by Ballmann, 1973, 1976):
* '' Apus wetmorei'', a swift.
* ''
Columba
Columba or Colmcille; gd, Calum Cille; gv, Colum Keeilley; non, Kolban or at least partly reinterpreted as (7 December 521 – 9 June 597 AD) was an Irish abbot and missionary evangelist credited with spreading Christianity in what is tod ...
omnisanctorum'' - one of the oldest
pigeon
Columbidae () is a bird family consisting of doves and pigeons. It is the only family in the order Columbiformes. These are stout-bodied birds with short necks and short slender bills that in some species feature fleshy ceres. They primarily ...
fossils known. It probably was more widespread and if so, the older name ''C. pisana'' would likely apply to it.
* ''Garganoaetus freudenthali'' and ''Garganoaetus murivorus'' - two species of falconid, the former larger than a
golden eagle
The golden eagle (''Aquila chrysaetos'') is a bird of prey living in the Northern Hemisphere. It is the most widely distributed species of eagle. Like all eagles, it belongs to the family Accipitridae. They are one of the best-known birds ...
, the latter well-sized; endemic. The smaller species, which likely is the stratigraphically oldest, is closely related to ''Aquila delphinensis'' from La Grive-Saint-Alban, France, according to Peter Ballmann in 1973. Its closest living relatives are the small eagles ('' Hieraaetus'', '' Spizaetus'', '' Lophaetus'').
* ''
Garganornis ballmanni
''Garganornis'' (meaning "Gargano bird") is an extinct genus of enormous flightless anatid waterfowl from the Late Miocene of Gargano, Italy. The genus contains one species, ''G. ballmanni'', named by Meijer in 2014. Its enormous size is thought ...
'', an extinct flightless giant goose-like waterfowl
* ''"Strix" perpasta'' - a true owl, perhaps the same as the widespread ''Bubo zeylonensis lamarmorae'', a
paleosubspecies
A chronospecies is a species derived from a sequential development pattern that involves continual and uniform changes from an extinct ancestral form on an evolutionary scale. The sequence of alterations eventually produces a population that is p ...
of the
brown fish-owl
The brown fish owl (''Ketupa zeylonensis'') is a fish owl species in the family (biology), family known as typical owls, Strigidae. It is native from Turkey to South Asia, South and Southeast Asia. Due its wide distribution it is listed as Least ...
(Mlíkovský 2002) but this
taxon
In biology, a taxon ( back-formation from '' taxonomy''; plural taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular n ...
was usually known from later times.
* '' Tyto'' - 2 or 3 species of barn-owls. The largest, '' T. gigantea'', was up to twice as massive as the living
eagle-owl
The American (North and South America) horned owls and the Old World eagle-owls make up the genus ''Bubo'', at least as traditionally described. The genus name ''Bubo'' is Latin for the Eurasian eagle-owl.
This genus contains 19 species that ar ...
'' Bubo bubo''. '' T. robusta'' was also large; this species and the former were endemic but actually seem to have been chronosubspecies. The supposed remains of the smaller ''T. sanctialbani'' found at Gargano are now placed in the widespread ''Tyto balearica''.
* an indeterminate woodpecker.
Languages
According to Pellegrini, Gargano is home t area IIIb of Southern Italo-Romance varieties. Each town, in turn, speaks its own sub-variety. Th Candeloro (or ''Candelaro'' in Italian) river defines the boundaries of the promontory as well as the borders with area IIb (that of Foggiano varieties).
* Butler, M., 1980. The giant erinaceid insectivore, Deinogalerix Freudenthal, from the upper Miocene of Gargano, Italy. Scripta Geologica 57, 1-72.
* Daams, R., Freudenthal, M. (1985): "''Stertomys laticrestatus'', a new glirid (dormice, Rodentia) from the insular fauna of Gargano (Prov. of Foggia, Italy)." '' Scripta Geologica'' 77: 21–27 (includes full text PDF)
* Freudenthal, M. (1972): "''Deinogalerix koenigswaldi'' nov. gen., nov. spec., a giant insectivore from the Neogene of Italy." '' Scripta Geologica'' 14: 1-1 includes full text PDF)
* Freudenthal, M. (1976): "Rodent stratigraphy of some Miocene fissure fillings in Gargano (prov. Foggia, Italy)". '' Scripta Geologica'' 37: 1-2 (includes full text PDF)
* Freudenthal, M. (1985) "Cricetidae (Rodentia) from the Neogene of Gargano (Prov. of Foggia, Italy)". '' Scripta Geologica'' 77: 29-76 (includes full text PDF)
* Freudenthal, M., Martín-Suárez, E. (2006): "Gliridae (Rodentia, Mammalia) from the Late Miocene Fissure Filling Biancone 1 (Gargano, Province of Foggia, Italy)." ''Palaeontologia Electronica'' 9.2.6A: 1-23.
* Leinders, J.J.M. (1984): "Hoplitomerycidae fam. nov. (Ruminantia, Mammalia) from Neogene fissure fillings in Gargano (Italy); part 1: The cranial osteology of Hoplitomeryx gen. nov. and a discussion on the classification of pecoran families". '' Scripta Geologica'' 70: 1-51, 9 plates.
*
* Mlíkovský, Jirí (2002): ''Cenozoic Birds of the World, Part 1: Europe'': 215. Ninox Press, Prague. PDF fulltext !-- This should be treated with extreme caution as regards merging of species. Splits are usually good though. See also critical review in Auk121:623-627 here http://www.findarticles.com/p/articles/mi_qa3793/is_200404/ai_n9396879 -->
* Parra, V.; Loreau, M. & Jaeger, J.-J. (1999): "Incisor size and community structure in rodents: two tests of the role of competition". ''Acta Oecologica'' 20(2): 93-101. (HTML abstract)
*
*
* (includes full text PDF)