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Tylomelania Centaurus Shell 2
''Tylomelania'' is a genus of freshwater snails which have an Operculum (gastropod), operculum, aquatic animal, aquatic gastropod mollusks in the family Pachychilidae. In the aquarium hobby, snails from this genus are commonly known as "rabbit snails" (not to be confused with Anaspidea, sea hares). A sister group (the closest relative) of ''Tylomelania'' is genus ''Pseudopotamis'' (2 species living on the Torres Strait Islands).von Rintelen T., Stelbrink B. Marwoto R. M. & Glaubrecht M. (2014). "A Snail Perspective on the Biogeography of Sulawesi, Indonesia: Origin and Intra-Island Dispersal of the Viviparous Freshwater Gastropod ''Tylomelania''". ''PLoS ONE'' 9(6): e98917. . ''Tylomelania'' and ''Pseudopotamis'' split in the Middle Miocene c. 19.5 Mya. Distribution These freshwater snails are Endemism, endemic to Sulawesi in Indonesia, with the vast majority restricted to Lake Poso and the Malili Lake system (consisting of Lake Matano, Matano and Lake Towuti, Towuti, and the ...
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Gastropod Shell
The gastropod shell is part of the body of a Gastropoda, gastropod or snail, a kind of mollusc. The shell is an exoskeleton, which protects from predators, mechanical damage, and dehydration, but also serves for muscle attachment and calcium storage. Some gastropods appear shell-less (slugs) but may have a remnant within the mantle, or in some cases the shell is reduced such that the body cannot be retracted within it (semi-slug). Some snails also possess an operculum that seals the opening of the shell, known as the Aperture (mollusc), aperture, which provides further protection. The study of mollusc shells is known as conchology. The biological study of gastropods, and other molluscs in general, is malacology. Shell morphology terms vary by species group. Shell layers The gastropod shell has three major layers secreted by the Mantle (mollusc), mantle. The calcareous central layer, tracum, is typically made of calcium carbonate precipitated into an organic matrix known as c ...
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Mollusk
Mollusca is the second-largest phylum of invertebrate animals after the Arthropoda, the members of which are known as molluscs or mollusks (). Around 85,000  extant species of molluscs are recognized. The number of fossil species is estimated between 60,000 and 100,000 additional species. The proportion of undescribed species is very high. Many taxa remain poorly studied. Molluscs are the largest marine phylum, comprising about 23% of all the named marine organisms. Numerous molluscs also live in freshwater and terrestrial habitats. They are highly diverse, not just in size and anatomical structure, but also in behaviour and habitat. The phylum is typically divided into 7 or 8  taxonomic classes, of which two are entirely extinct. Cephalopod molluscs, such as squid, cuttlefish, and octopuses, are among the most neurologically advanced of all invertebrates—and either the giant squid or the colossal squid is the largest known invertebrate species. The gas ...
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Lake Lontoa
A lake is an area filled with water, localized in a basin, surrounded by land, and distinct from any river or other outlet that serves to feed or drain the lake. Lakes lie on land and are not part of the ocean, although, like the much larger oceans, they do form part of the Earth's water cycle. Lakes are distinct from lagoons, which are generally coastal parts of the ocean. Lakes are typically larger and deeper than ponds, which also lie on land, though there are no official or scientific definitions. Lakes can be contrasted with rivers or streams, which usually flow in a channel on land. Most lakes are fed and drained by rivers and streams. Natural lakes are generally found in mountainous areas, rift zones, and areas with ongoing glaciation. Other lakes are found in endorheic basins or along the courses of mature rivers, where a river channel has widened into a basin. Some parts of the world have many lakes formed by the chaotic drainage patterns left over from the last ic ...
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Lake Towuti
Lake Towuti () is a lake in East Luwu Regency, South Sulawesi province, Indonesia. Surrounded by mountains, it is the largest lake of the island of Sulawesi and one of the five lakes of the Malili Lake system (the other being Matano, and the small Mahalona, Masapi and Lontoa (Wawantoa)).Gray, S.M., and J.S. McKinnon (2006). ''A comparative description of mating behaviour in the endemic telmatherinid fishes of Sulawesi's Malili Lakes.'' Environmental Biology of Fishes 75: 471–482 A river flows from the lake to the Boni Bay. The town Laronda is located on its shore. Ecology Together with the other lakes in the Malili Lake system, Towuti has a large number of endemic fishes ( Telmatherinid sail-fin silversides, ''Glossogobius'' and '' Mugilogobius'' gobies, ''Nomorhamphus'' halfbeaks and ''Oryzias'' ricefishes), ''Caridina'' shrimps, Parathelphusid crabs ('' Nautilothelphusa'', '' Parathelphusa'' and '' Syntripsa'') and ''Tylomelania'' snails. Among these, ''Caridina spong ...
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Lake Matano
Lake Matano ( id, Danau Matano), also known as Matana, is a natural lake in East Luwu Regency, South Sulawesi province, Indonesia. With a depth of , it is the deepest lake in Indonesia (ranked by maximum depth), the 10th deepest lake in the world and the deepest lake on an island by maximum depth. The surface elevation from mean sea level is only , which means that the deepest portion of the lake is below sea level (cryptodepression). It is one of the two major lakes (the other being Lake Towuti) in the Malili Lake system. Endemic animals and plants Lake Matano is home to many species of endemic fish and other animals (e.g. ''Caridina'' shrimps, Parathelphusid crabs and ''Tylomelania'' snails) as well as many plants. The endemic fishes of Matano have been compared to the species swarms of the Rift Valley Lakes of Africa. While not as diverse, they are thought to have all arisen from a single ancestor species and diversified into numerous different species, which now fill ma ...
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Lake Poso
Lake Poso ( id, Danau Poso) is a lake in Central Sulawesi, Indonesia, and the third-deepest lake in Indonesia. The town of Pendolo is situated at the southern end of the lake, the town of Tentena is located at the northern end, while a number of smaller villages dot the shoreline. The lake drains into the Poso River at Tentena, which flows into the Molucca Sea at the town of Poso. Ecology The lake contains various fish, including the eel ''Anguilla marmorata'' which migrates between the lake and the sea, and 11 fish species that are endemic to the lake, notably buntingi ricefish ('' Adrianichthys'', ''Oryzias nebulosus'', '' O. nigrimas'' and '' O. orthognathus''), gobies (''Mugilogobius amadi'' and '' M. sarasinorum''), and the halfbeak ''Nomorhamphus celebensis''. These endemics are all highly threatened; in some cases possibly already extinct. One of the reasons for the drastic decline of the native fish are introduced, non-native species, particularly Mozambique t ...
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Indonesia
Indonesia, officially the Republic of Indonesia, is a country in Southeast Asia and Oceania between the Indian and Pacific oceans. It consists of over 17,000 islands, including Sumatra, Java, Sulawesi, and parts of Borneo and New Guinea. Indonesia is the world's largest archipelagic state and the 14th-largest country by area, at . With over 275 million people, Indonesia is the world's fourth-most populous country and the most populous Muslim-majority country. Java, the world's most populous island, is home to more than half of the country's population. Indonesia is a presidential republic with an elected legislature. It has 38 provinces, of which nine have special status. The country's capital, Jakarta, is the world's second-most populous urban area. Indonesia shares land borders with Papua New Guinea, East Timor, and the eastern part of Malaysia, as well as maritime borders with Singapore, Vietnam, Thailand, the Philippines, Australia, Palau, and India ...
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Sulawesi
Sulawesi (), also known as Celebes (), is an island in Indonesia. One of the four Greater Sunda Islands, and the world's eleventh-largest island, it is situated east of Borneo, west of the Maluku Islands, and south of Mindanao and the Sulu Archipelago. Within Indonesia, only Sumatra, Borneo, and New Guinea, Papua are larger in territory, and only Java and Sumatra have larger populations. The landmass of Sulawesi includes four peninsulas: the northern Minahassa Peninsula, Minahasa Peninsula, the East Peninsula, Sulawesi, East Peninsula, the South Peninsula, Sulawesi, South Peninsula, and the Southeast Peninsula, Sulawesi, Southeast Peninsula. Three gulfs separate these peninsulas: the Gulf of Tomini between the northern Minahasa and East peninsulas, the Tolo Gulf between the East and Southeast peninsulas, and the Bone Gulf between the South and Southeast peninsulas. The Strait of Makassar runs along the western side of the island and separates the island from Borneo. Etymology ...
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Endemism
Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found elsewhere. For example, the Cape sugarbird is found exclusively in southwestern South Africa and is therefore said to be ''endemic'' to that particular part of the world. An endemic species can be also be referred to as an ''endemism'' or in scientific literature as an ''endemite''. For example '' Cytisus aeolicus'' is an endemite of the Italian flora. '' Adzharia renschi'' was once believed to be an endemite of the Caucasus, but it was later discovered to be a non-indigenous species from South America belonging to a different genus. The extreme opposite of an endemic species is one with a cosmopolitan distribution, having a global or widespread range. A rare alternative term for a species that is endemic is "precinctive", which applies to ...
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Tylomelania Map
''Tylomelania'' is a genus of freshwater snails which have an operculum, aquatic gastropod mollusks in the family Pachychilidae. In the aquarium hobby, snails from this genus are commonly known as "rabbit snails" (not to be confused with sea hares). A sister group (the closest relative) of ''Tylomelania'' is genus ''Pseudopotamis'' (2 species living on the Torres Strait Islands).von Rintelen T., Stelbrink B. Marwoto R. M. & Glaubrecht M. (2014). "A Snail Perspective on the Biogeography of Sulawesi, Indonesia: Origin and Intra-Island Dispersal of the Viviparous Freshwater Gastropod ''Tylomelania''". ''PLoS ONE'' 9(6): e98917. . ''Tylomelania'' and ''Pseudopotamis'' split in the Middle Miocene c. 19.5 Mya. Distribution These freshwater snails are endemic to Sulawesi in Indonesia, with the vast majority restricted to Lake Poso and the Malili Lake system (consisting of Matano and Towuti, and the smaller Lontoa (Wawantoa), Mahalona, Masapi).von Rintelen T., von Rintele ...
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Middle Miocene
The Middle Miocene is a sub-epoch of the Miocene Epoch made up of two stages: the Langhian and Serravallian stages. The Middle Miocene is preceded by the Early Miocene. The sub-epoch lasted from 15.97 ± 0.05 Ma to 11.608 ± 0.005 Ma (million years ago). During this period, a sharp drop in global temperatures took place. This event is known as the Middle Miocene Climate Transition. For the purpose of establishing European Land Mammal Ages The European Land Mammal Mega Zones (abbreviation: ELMMZ, more commonly known as European land mammal ages or ELMA) are zones in rock layers that have a specific assemblage of fossils (biozones) based on occurrences of fossil assemblages of Europe ... this sub-epoch is equivalent to the Astaracian age. External links GeoWhen Database - Middle Miocene .02 02 * * {{geochronology-stub ...
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Torres Strait Islands
The Torres Strait Islands are a group of at least 274 small islands in the Torres Strait, a waterway separating far northern continental Australia's Cape York Peninsula and the island of New Guinea. They span an area of , but their total land area is . The Islands have been inhabited by the indigenous Torres Strait Islanders. Lieutenant James Cook first claimed British sovereignty over the eastern part of Australia at Possession Island, Queensland, Possession Island in 1770, but British administrative control only began in the Torres Strait Islands in 1862. The islands are now mostly part of Queensland, a constituent State of the Australia, Commonwealth of Australia, but are administered by the Torres Strait Regional Authority, a statutory authority of the Australian federal government. A few islands very close to the coast of mainland New Guinea belong to the Western Province (Papua New Guinea), Western Province of Papua New Guinea, most importantly Daru Island with the provin ...
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