Tropidostoma Dubium
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Tropidostoma Dubium
''Tropidostoma'' is a medium-sized herbivorous oudenodontid dicynodont therapsid that lived during the Late Permian (Lopingian) period in South Africa. The first ''Tropidostoma'' fossil was described by Harry Govier Seeley in 1889. Later two subspecies were identified. ''Tropidostoma'' fossils are an index fossil in a biozone of the Karoo Basin known as the ''Tropidostoma'' Assemblage Zone. This biozone is characterized by the presence of this species in association with another dicynodont species, '' Endothiodon uniseries''. History of discovery The first ''Tropidostoma'' fossil material was found during a field expedition in the Upper Permian-aged Teekloof Formation of the Beaufort Group. This material was later described by Seeley (1889) in a study in which he described two fossils which had been named ''Dicynodon microtrema'' and ''Tropidostoma dunni''. In 1915, several years after Seeley's death, the paleontologist, Robert Broom, reexamined the same material and discovered ...
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Late Permian
Late may refer to: * LATE, an acronym which could stand for: ** Limbic-predominant age-related TDP-43 encephalopathy, a proposed form of dementia ** Local-authority trading enterprise, a New Zealand business law ** Local average treatment effect, a concept in econometrics Music * Late (album), ''Late'' (album), a 2000 album by The 77s * Late!, a pseudonym used by Dave Grohl on his ''Pocketwatch (album), Pocketwatch'' album * Late (rapper), an underground rapper from Wolverhampton * Late (song), "Late" (song), a song by Blue Angel * "Late", a song by Kanye West from ''Late Registration'' Other * Late (Tonga), an uninhabited volcanic island southwest of Vavau in the kingdom of Tonga * Late (The Handmaid's Tale), "Late" (''The Handmaid's Tale''), a television episode * LaTe, Laivateollisuus, Oy Laivateollisuus Ab, a defunct shipbuilding company * Late may refer to a person who is Dead See also

* * * ''Lates'', a genus of fish in the lates perch family * Later (other) ...
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Robert Broom
Robert Broom FRS FRSE (30 November 1866 6 April 1951) was a British- South African doctor and palaeontologist. He qualified as a medical practitioner in 1895 and received his DSc in 1905 from the University of Glasgow. From 1903 to 1910, he was professor of zoology and geology at Victoria College, Stellenbosch, South Africa, and subsequently he became keeper of vertebrate palaeontology at the South African Museum, Cape Town. Life Broom was born at 66 Back Sneddon Street in Paisley, Renfrewshire, Scotland, the son of John Broom, a designer of calico prints and Paisley shawls, and Agnes Hunter Shearer. In 1893, he married Mary Baird Baillie, his childhood sweetheart. In his medical studies at the University of Glasgow Broom specialised in obstetrics. After graduating in 1895 he travelled to Australia, supporting himself by practising medicine. He settled in South Africa in 1897, just prior to the South African War. From 1903 to 1910, he was professor of Zoology and Geolog ...
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Dicynodonts
Dicynodontia is an extinct clade of anomodonts, an extinct type of non-mammalian therapsid. Dicynodonts were Herbivore, herbivorous animals with a pair of tusks, hence their name, which means 'two dog tooth'. Members of the group possessed a horny, typically toothless beak, unique amongst all synapsids. Dicynodonts first appeared in Southern Pangaea during the Guadalupian, mid-Permian, ca. 270–260 million years ago, and became globally distributed and the dominant herbivorous animals in the Permian, Late Permian, ca. 260–252 Mya. They were devastated by the Permian–Triassic extinction event, end-Permian Extinction that wiped out most other therapsids ca. 252 Mya. They rebounded during the Triassic but died out towards the end of that period. They were the most successful and diverse of the non-mammalian therapsids, with over 70 genus, genera known, varying from rat-sized Fossorial, burrowers to elephant-sized Browsing (herbivory), browsers. Characteristics The dicynodont s ...
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List Of Therapsids
This list of therapsids is an attempt to create a comprehensive listing of all genera that have ever been included in the Therapsida excluding mammals and purely vernacular terms. The list includes all commonly accepted genera, but also genera that are now considered invalid, doubtful ('' nomina dubia''), or were not formally published (''nomina nuda''), as well as junior synonyms of more established names, and genera that are no longer considered therapsids. The list currently contains 510 generic names. Naming conventions and terminology Naming conventions and terminology follow the International Code of Zoological Nomenclature. Technical terms used include: * Junior synonym: A name which describes the same taxon as a previously published name. If two or more genera are formally designated and the type specimens are later assigned to the same genus, the first to be published (in chronological order) is the senior synonym, and all other instances are junior synonyms. Senior synon ...
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Bulbasaurus
''Bulbasaurus'' (meaning "bulbous reptile") is an extinct genus of dicynodont that is known from the Lopingian epoch of the Late Permian period of what is now South Africa, containing the type and only species ''B. phylloxyron''. It was formerly considered as belonging to ''Tropidostoma''; however, due to numerous differences from ''Tropidostoma'' in terms of skull morphology and size, it has been reclassified the earliest known member of the family Geikiidae, and the only member of the group known from the ''Tropidostoma'' Assemblage Zone. Within the Geikiidae, it has been placed close to '' Aulacocephalodon'', although a more basal position is not implausible. ''Bulbasaurus'' was ostensibly not directly named after the Pokémon Bulbasaur, but rather after its nasal bosses, which are unusually bulbous among geikiids; however, the describers noted that the similarity in name "may not be entirely coincidental." Additionally, the specific name of the type species means "leaf raz ...
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Oudenodon
''Oudenodon'' is an extinct genus of dicynodont. It was common throughout southern Africa during the Late Permian. Several species of ''Oudenodon'' are known. Both ''O. bainii'', the type species, and ''O. grandis'' are known from South Africa. Specimens of ''O. luangwensis'' have been found from Zambia. One species, ''O. sakamenensis'', is the only Permian therapsid yet known from Madagascar.Mazin, J. M. and King, G. M. (1991). The first dicynodont from the Late Permian of Malagasy. ''Palaeontology'' 34:837–842. It is the type genus of the family Oudenodontidae, which includes members such as '' Tropidostoma''. See also * List of therapsids References The main groups of non-mammalian synapsids at Mikko's Phylogeny Archive External links ''Oudenodon''in the Paleobiology Database The Paleobiology Database is an online resource for information on the distribution and classification of fossil animals, plants, and microorganisms. History The Paleobiology Database (PBDB) or ...
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Keratin
Keratin () is one of a family of structural fibrous proteins also known as ''scleroproteins''. Alpha-keratin (α-keratin) is a type of keratin found in vertebrates. It is the key structural material making up scales, hair, nails, feathers, horns, claws, hooves, and the outer layer of skin among vertebrates. Keratin also protects epithelial cells from damage or stress. Keratin is extremely insoluble in water and organic solvents. Keratin monomers assemble into bundles to form intermediate filaments, which are tough and form strong unmineralized epidermal appendages found in reptiles, birds, amphibians, and mammals. Excessive keratinization participate in fortification of certain tissues such as in horns of cattle and rhinos, and armadillos' osteoderm. The only other biological matter known to approximate the toughness of keratinized tissue is chitin. Keratin comes in two types, the primitive, softer forms found in all vertebrates and harder, derived forms found only amon ...
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Maxilla
The maxilla (plural: ''maxillae'' ) in vertebrates is the upper fixed (not fixed in Neopterygii) bone of the jaw formed from the fusion of two maxillary bones. In humans, the upper jaw includes the hard palate in the front of the mouth. The two maxillary bones are fused at the intermaxillary suture, forming the anterior nasal spine. This is similar to the mandible (lower jaw), which is also a fusion of two mandibular bones at the mandibular symphysis. The mandible is the movable part of the jaw. Structure In humans, the maxilla consists of: * The body of the maxilla * Four processes ** the zygomatic process ** the frontal process of maxilla ** the alveolar process ** the palatine process * three surfaces – anterior, posterior, medial * the Infraorbital foramen * the maxillary sinus * the incisive foramen Articulations Each maxilla articulates with nine bones: * two of the cranium: the frontal and ethmoid * seven of the face: the nasal, zygomatic, lacrimal, inferior n ...
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Dentition
Dentition pertains to the development of teeth and their arrangement in the mouth. In particular, it is the characteristic arrangement, kind, and number of teeth in a given species at a given age. That is, the number, type, and morpho-physiology (that is, the relationship between the shape and form of the tooth in question and its inferred function) of the teeth of an animal. Animals whose teeth are all of the same type, such as most non-mammalian vertebrates, are said to have '' homodont'' dentition, whereas those whose teeth differ morphologically are said to have '' heterodont'' dentition. The dentition of animals with two successions of teeth (deciduous, permanent) is referred to as ''diphyodont'', while the dentition of animals with only one set of teeth throughout life is ''monophyodont''. The dentition of animals in which the teeth are continuously discarded and replaced throughout life is termed ''polyphyodont''. The dentition of animals in which the teeth are set in so ...
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Bidentalia
Bidentalia is a group of dicynodont therapsids. Bidentalia was one of the first names used to describe dicynodonts; the group was established in 1876, while the name "bidentals" dates back as far as 1845. With the increasing prominence of phylogenetics, the group was redefined as a clade in 2009. Bidentalia is now considered a stem-based taxon that includes all taxa more closely related to '' Aulacephalodon bainii'' and ''Dicynodon lacerticeps'' than '' Emydops arctatus''. History In 1845, South African geologist Andrew Geddes Bain described the first known dicynodonts as "bidentals" for their two prominent tusks. "Bidental" was the first name ever used for a group of non-mammalian synapsids. The name Dicynodontia, which is more commonly used to refer to these animals, was erected by English paleontologist Richard Owen in 1860. Realizing that Bain's "bidentals" predated his "dicynodonts", Owen named Bidentalia in 1876 as a replacement name for Dicynodontia. Owen described Bident ...
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Clade
A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English term, the equivalent Latin term ''cladus'' (plural ''cladi'') is often used in taxonomical literature. The common ancestor may be an individual, a population, or a species (extinct or extant). Clades are nested, one in another, as each branch in turn splits into smaller branches. These splits reflect evolutionary history as populations diverged and evolved independently. Clades are termed monophyletic (Greek: "one clan") groups. Over the last few decades, the cladistic approach has revolutionized biological classification and revealed surprising evolutionary relationships among organisms. Increasingly, taxonomists try to avoid naming taxa that are not clades; that is, taxa that are not monophyletic. Some of the relationships between organisms ...
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Sexual Dimorphism
Sexual dimorphism is the condition where the sexes of the same animal and/or plant species exhibit different morphological characteristics, particularly characteristics not directly involved in reproduction. The condition occurs in most animals and some plants. Differences may include secondary sex characteristics, size, weight, colour, markings, or behavioural or cognitive traits. These differences may be subtle or exaggerated and may be subjected to sexual selection and natural selection. The opposite of dimorphism is ''monomorphism'', which is when both biological sexes are phenotypically indistinguishable from each other. Overview Ornamentation and coloration Common and easily identified types of dimorphism consist of ornamentation and coloration, though not always apparent. A difference in coloration of sexes within a given species is called sexual dichromatism, which is commonly seen in many species of birds and reptiles. Sexual selection leads to the exaggerated dim ...
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