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Trimmatothele
''Trimmatothele'' is a genus of saxicolous (rock-dwelling), crustose lichens in the family Verrucariaceae. It has five species. The genus was formally published by lichenologist Alexander Zahlbruckner in 1903. The type species, '' Trimmatothele perquisita'', was originally collected from Norway by Johannes M. Norman, who placed it in the genus ''Coniothele'' in 1868. Diagnostic characteristics of ''Trimmatothele'' include a thin thallus with a smooth surface; small perithecia that are partially immersed in the substrata and have an involucrellum; asci that contain multiple ascospores; and small, simple ascospores (i.e., without any septa). ''Trimmatothele'' has been described as one of the most poorly known genera of lichens due to the rarity of its species, the few available herbarium specimens, and some missing type specimens In biology, a type is a particular specimen (or in some cases a group of specimens) of an organism to which the scientific name of that organism is ...
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Trimmatothele Glacialis
''Trimmatothele'' is a genus of saxicolous (rock-dwelling), crustose lichens in the family Verrucariaceae. It has five species. The genus was formally published by lichenologist Alexander Zahlbruckner in 1903. The type species, '' Trimmatothele perquisita'', was originally collected from Norway by Johannes M. Norman, who placed it in the genus ''Coniothele'' in 1868. Diagnostic characteristics of ''Trimmatothele'' include a thin thallus with a smooth surface; small perithecia that are partially immersed in the substrata and have an involucrellum; asci that contain multiple ascospores; and small, simple ascospores (i.e., without any septa). ''Trimmatothele'' has been described as one of the most poorly known genera of lichens due to the rarity of its species, the few available herbarium specimens, and some missing type specimens In biology, a type is a particular specimen (or in some cases a group of specimens) of an organism to which the scientific name of that organism is ...
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Trimmatothele Maritima
''Trimmatothele'' is a genus of saxicolous (rock-dwelling), crustose lichens in the family Verrucariaceae. It has five species. The genus was formally published by lichenologist Alexander Zahlbruckner in 1903. The type species, '' Trimmatothele perquisita'', was originally collected from Norway by Johannes M. Norman, who placed it in the genus ''Coniothele'' in 1868. Diagnostic characteristics of ''Trimmatothele'' include a thin thallus with a smooth surface; small perithecia that are partially immersed in the substrata and have an involucrellum; asci that contain multiple ascospores; and small, simple ascospores (i.e., without any septa). ''Trimmatothele'' has been described as one of the most poorly known genera of lichens due to the rarity of its species, the few available herbarium specimens, and some missing type specimens. Species *''Trimmatothele glacialis ''Trimmatothele'' is a genus of saxicolous (rock-dwelling), crustose lichens in the family Verrucariaceae. It has ...
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Trimmatothele Perquisita
''Trimmatothele'' is a genus of saxicolous (rock-dwelling), crustose lichens in the family Verrucariaceae. It has five species. The genus was formally published by lichenologist Alexander Zahlbruckner in 1903. The type species, '' Trimmatothele perquisita'', was originally collected from Norway by Johannes M. Norman, who placed it in the genus ''Coniothele'' in 1868. Diagnostic characteristics of ''Trimmatothele'' include a thin thallus with a smooth surface; small perithecia that are partially immersed in the substrata and have an involucrellum; asci that contain multiple ascospores; and small, simple ascospores (i.e., without any septa). ''Trimmatothele'' has been described as one of the most poorly known genera of lichens due to the rarity of its species, the few available herbarium specimens, and some missing type specimens. Species *''Trimmatothele glacialis'' *''Trimmatothele maritima ''Trimmatothele'' is a genus of saxicolous (rock-dwelling), crustose lichens in the ...
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Trimmatothele Petri
''Trimmatothele'' is a genus of saxicolous (rock-dwelling), crustose lichens in the family Verrucariaceae. It has five species. The genus was formally published by lichenologist Alexander Zahlbruckner in 1903. The type species, ''Trimmatothele perquisita'', was originally collected from Norway by Johannes M. Norman, who placed it in the genus ''Coniothele'' in 1868. Diagnostic characteristics of ''Trimmatothele'' include a thin thallus with a smooth surface; small perithecia that are partially immersed in the substrata and have an involucrellum; asci that contain multiple ascospores; and small, simple ascospores (i.e., without any septa). ''Trimmatothele'' has been described as one of the most poorly known genera of lichens due to the rarity of its species, the few available herbarium specimens, and some missing type specimens. Species *''Trimmatothele glacialis'' *''Trimmatothele maritima'' *''Trimmatothele perquisita ''Trimmatothele'' is a genus of saxicolous (rock-dwell ...
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Trimmatothele Umbellulariae
''Trimmatothele'' is a genus of saxicolous (rock-dwelling), crustose lichens in the family Verrucariaceae. It has five species. The genus was formally published by lichenologist Alexander Zahlbruckner in 1903. The type species, ''Trimmatothele perquisita'', was originally collected from Norway by Johannes M. Norman, who placed it in the genus ''Coniothele'' in 1868. Diagnostic characteristics of ''Trimmatothele'' include a thin thallus with a smooth surface; small perithecia that are partially immersed in the substrata and have an involucrellum; asci that contain multiple ascospores; and small, simple ascospores (i.e., without any septa). ''Trimmatothele'' has been described as one of the most poorly known genera of lichens due to the rarity of its species, the few available herbarium specimens, and some missing type specimens. Species *''Trimmatothele glacialis'' *''Trimmatothele maritima'' *''Trimmatothele perquisita'' *''Trimmatothele petri ''Trimmatothele'' is a genus ...
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Verrucariaceae
The Verrucariaceae are a family of mostly lichenised fungi in the order Verrucariales. The lichen-forming species, which comprise the vast majority of the family, have a wide variety of thallus forms, and include crustose (crust-like), foliose (bushy), and squamulose (scaly) representatives. Several characteristics of the spore-bearing structures, the ascomata, define the family, including their perithecioid form–more or less spherical or flask-shaped, with a single opening and otherwise completely enclosed by a wall. Squamulose members of the Verrucariaceae with simple ascospores (lacking partitions called septa), and without algae in the spore-bearing region are known as lichens; there are more than 80 of these species. The family has several dozen lichenicolous (lichen-dwelling) examples, including a few genera that contain solely lichenicolous members. An unusually diverse variety of photobiont partners have been recorded, mostly green algae, but also brown algae and ...
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Johannes M
Johannes is a Medieval Latin form of the personal name that usually appears as "John" in English language contexts. It is a variant of the Greek and Classical Latin variants (Ιωάννης, ''Ioannes''), itself derived from the Hebrew name '' Yehochanan'', meaning "Yahweh is gracious". The name became popular in Northern Europe, especially in Germany because of Christianity. Common German variants for Johannes are ''Johann'', ''Hannes'', '' Hans'' (diminutized to ''Hänschen'' or ''Hänsel'', as known from "''Hansel and Gretel''", a fairy tale by the Grimm brothers), '' Jens'' (from Danish) and ''Jan'' (from Dutch, and found in many countries). In the Netherlands, Johannes was without interruption the most common masculine birth name until 1989. The English equivalent for Johannes is John. In other languages *Joan, Jan, Gjon, Gjin and Gjovalin in Albanian *'' Yoe'' or '' Yohe'', uncommon American form''Dictionary of American Family Names'', Oxford University Press, 2013. *Yaḥy ...
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Ascospore
An ascus (; ) is the sexual spore-bearing cell produced in ascomycete fungi. Each ascus usually contains eight ascospores (or octad), produced by meiosis followed, in most species, by a mitotic cell division. However, asci in some genera or species can occur in numbers of one (e.g. ''Monosporascus cannonballus''), two, four, or multiples of four. In a few cases, the ascospores can bud off conidia that may fill the asci (e.g. ''Tympanis'') with hundreds of conidia, or the ascospores may fragment, e.g. some ''Cordyceps'', also filling the asci with smaller cells. Ascospores are nonmotile, usually single celled, but not infrequently may be coenocytic (lacking a septum), and in some cases coenocytic in multiple planes. Mitotic divisions within the developing spores populate each resulting cell in septate ascospores with nuclei. The term ocular chamber, or oculus, refers to the epiplasm (the portion of cytoplasm not used in ascospore formation) that is surrounded by the "bourrelet ...
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Lichen Genera
A lichen ( , ) is a composite organism that arises from algae or cyanobacteria living among filaments of multiple fungi species in a mutualistic relationship.Introduction to Lichens – An Alliance between Kingdoms
. University of California Museum of Paleontology.
Lichens have properties different from those of their component organisms. They come in many colors, sizes, and forms and are sometimes plant-like, but are not s. They may have tiny, leafless branches (); flat leaf-like structures (

Eurotiomycetes Genera
Eurotiomycetes is a large class of ascomycetes with cleistothecial ascocarps within the subphylum Pezizomycotina, currently containing around 3810 species according to the Catalogue of Life. It is the third largest lichenized class, with more than 1200 lichen species that are mostly bitunicate in the formation of asci. It contains most of the fungi previously known morphologically as "Plectomycetes". Systematics and phylogeny Internal relationships The class Eurotiomycetes was circumscribed in 1997 by Swedish mycologists Ove Erik Eriksson and Katarina Winka. At that time it only contained the order Eurotiales, which together with the next order added, Onygenales, form a monophyletic group comprising most of the fungi in "Plectomycetes", a group no longer in use that unified fungi under exclusively morphological characteristics. As more orders were added to Eurotiomycetes, the first two along with Arachnomycetales became constrained to the first subclass, Eurotiomycetidae. In 2 ...
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Verrucariales
Verrucariales is an order of ascomycetous fungi within the subclass Chaetothyriomycetidae of the class Eurotiomycetes. Although most of the Verrucariales are lichenised, the family Sarcopyreniaceae consists of 11 species of lichenicolous (lichen-dwelling) fungi. Phylogenomic analysis suggests that the divergence between the lichenized Verrucariales and nonlichenized Chaetothyriales The Chaetothyriales are an order of ascomycetous fungi in the class Eurotiomycetes and within the subclass Chaetothyriomycetidae. The order was circumscribed in 1987 by mycologist Margaret Elizabeth Barr-Bigelow. Families and genera , Species ... occurred about 131 million years ago. Genera of uncertain placement There are some genera in the Verrucariales that have not been placed with certainty into any family. These are: *'' Botryolepraria'' – 2 spp. *'' Gemmaspora'' – 1 sp. *'' Kalbiana'' – 1 sp. *'' Merismatium'' – 10 spp. References Ascomycota orders Lichen orders ...
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Type (biology)
In biology, a type is a particular specimen (or in some cases a group of specimens) of an organism to which the scientific name of that organism is formally attached. In other words, a type is an example that serves to anchor or centralizes the defining features of that particular taxon. In older usage (pre-1900 in botany), a type was a taxon rather than a specimen. A taxon is a scientifically named grouping of organisms with other like organisms, a set that includes some organisms and excludes others, based on a detailed published description (for example a species description) and on the provision of type material, which is usually available to scientists for examination in a major museum research collection, or similar institution. Type specimen According to a precise set of rules laid down in the International Code of Zoological Nomenclature (ICZN) and the International Code of Nomenclature for algae, fungi, and plants (ICN), the scientific name of every taxon is almost al ...
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