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Verrucariales
Verrucariales is an order of ascomycetous fungi within the subclass Chaetothyriomycetidae of the class Eurotiomycetes. Although most of the Verrucariales are lichenised, the family Sarcopyreniaceae consists of 11 species of lichenicolous (lichen-dwelling) fungi. Phylogenomic analysis suggests that the divergence between the lichenized Verrucariales and nonlichenized Chaetothyriales The Chaetothyriales are an order of ascomycetous fungi in the class Eurotiomycetes and within the subclass Chaetothyriomycetidae. The order was circumscribed in 1987 by mycologist Margaret Elizabeth Barr-Bigelow. Families and genera , Species ... occurred about 131 million years ago. Genera of uncertain placement There are some genera in the Verrucariales that have not been placed with certainty into any family. These are: *'' Botryolepraria'' – 2 spp. *'' Gemmaspora'' – 1 sp. *'' Kalbiana'' – 1 sp. *'' Merismatium'' – 10 spp. References Ascomycota orders Lichen orders ...
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Verrucariaceae
The Verrucariaceae are a family of mostly lichenised fungi in the order Verrucariales. The lichen-forming species, which comprise the vast majority of the family, have a wide variety of thallus forms, and include crustose (crust-like), foliose (bushy), and squamulose (scaly) representatives. Several characteristics of the spore-bearing structures, the ascomata, define the family, including their perithecioid form–more or less spherical or flask-shaped, with a single opening and otherwise completely enclosed by a wall. Squamulose members of the Verrucariaceae with simple ascospores (lacking partitions called septa), and without algae in the spore-bearing region are known as lichens; there are more than 80 of these species. The family has several dozen lichenicolous (lichen-dwelling) examples, including a few genera that contain solely lichenicolous members. An unusually diverse variety of photobiont partners have been recorded, mostly green algae, but also brown algae and ...
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Verrucariales
Verrucariales is an order of ascomycetous fungi within the subclass Chaetothyriomycetidae of the class Eurotiomycetes. Although most of the Verrucariales are lichenised, the family Sarcopyreniaceae consists of 11 species of lichenicolous (lichen-dwelling) fungi. Phylogenomic analysis suggests that the divergence between the lichenized Verrucariales and nonlichenized Chaetothyriales The Chaetothyriales are an order of ascomycetous fungi in the class Eurotiomycetes and within the subclass Chaetothyriomycetidae. The order was circumscribed in 1987 by mycologist Margaret Elizabeth Barr-Bigelow. Families and genera , Species ... occurred about 131 million years ago. Genera of uncertain placement There are some genera in the Verrucariales that have not been placed with certainty into any family. These are: *'' Botryolepraria'' – 2 spp. *'' Gemmaspora'' – 1 sp. *'' Kalbiana'' – 1 sp. *'' Merismatium'' – 10 spp. References Ascomycota orders Lichen orders ...
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Eurotiomycetes
Eurotiomycetes is a large class of ascomycetes with cleistothecial ascocarps within the subphylum Pezizomycotina, currently containing around 3810 species according to the Catalogue of Life. It is the third largest lichen A lichen ( , ) is a composite organism that arises from algae or cyanobacteria living among filaments of multiple fungi species in a mutualistic relationship. It contains most of the fungi previously known morphologically as "Plectomycetes".


Systematics and phylogeny


Internal relationships

The class Eurotiomycetes was circumscription (taxonomy), circumscribed in 1997 by Sweden, Swedish mycologists Ove Erik Eriksson and Katarina Winka. At that time ...

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Adelococcaceae
The Adelococcaceae are a family of fungi in the order Verrucariales. Species are mostly found in north temperate regions, and are biotrophic or necrotrophic A fungus ( : fungi or funguses) is any member of the group of eukaryotic organisms that includes microorganisms such as yeasts and molds, as well as the more familiar mushrooms. These organisms are classified as a kingdom, separately from t ... on lichens. The family was proposed by mycologist Dagmar Triebel in 1993. Genera * '' Adelococcus'' – 4 spp. * '' Pseudopyrenidium'' – 2 spp. * '' Sagediopsis'' – 11 spp. References Verrucariales Ascomycota families Taxa described in 1993 Lichenicolous fungi Taxa named by Dagmar Triebel {{Eurotiomycetes-stub ...
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Botryolepraria
''Botryolepraria'' is a genus of saxicolous (rock-dwelling), byssoid (cottony) lichens of uncertain familial placement in the order Verrucariales. It has two species. Both species grow in damp conditions, such as on cave walls, particularly in areas with minimal light. Taxonomy The genus was circumscribed by Antonio Canals, Mariona Hernández-Mariné, Antonio Gómez-Bolea, and Xavier Llimona in 1997, as a segregate of genus ''Lepraria'', with the widespread and common lichen '' B. lesdainii'' as the type, and at that time, only species. The type specimen was collected by French lichenologist Maurice Bouly de Lesdain from a wall in Les Baraques (Calais, France). The genus name combines the Greek-derived ''botryon'' ("cluster of berries", referring to the microscopic shrub-like clusters of fungal hyphae and spherical algal cells) with its namesake genus, ''Lepraria''. Although some later authors did not accept the proposed genus as different from ''Lepraria'', later molecul ...
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Merismatium
''Merismatium'' is a genus of lichenicolous (lichen-dwelling) fungi of uncertain familial placement in the order Verrucariales. The genus was circumscribed in 1898 by Friedrich Wilhelm Zopf. Species , Species Fungorum (in the Catalogue of Life) accepts six species of ''Merismatium''. Host information is from Diederich et al. (2018). * '' Merismatium coccotremicola'' – host: '' Coccotrema'' * '' Merismatium corae'' – host: '' Cora'' * '' Merismatium deminutum'' – hosts: '' Polyblastia'', ''Staurothele'', '' Thelidium'', '' Verrucaria'' * '' Merismatium nigritellum'' – hosts: '' Catapyrenium'', ''Leptogium'', ''Lichenomphalia'', ''Lopadium'', '' Micarea'', '' Mycobilimbia'', ''Ochrolechia'', ''Protoblastenia'' * '' Merismatium peregrinum'' – host: ''Rimularia'' * '' Merismatium thamnoliicola'' – host: ''Thamnolia ''Thamnolia'' is a genus of lichens in the family Icmadophilaceae. Members of the genus are commonly called whiteworm lichens. Two species of ''T ...
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Sarcopyreniaceae
''Sarcopyrenia'' is a genus of lichenicolous (lichen-dwelling) fungi. It has 11 species. It is the only genus in Sarcopyreniaceae, a family in the order Verrucariales. ''Sarcopyrenia'' was circumscribed by Finnish lichenologist William Nylander in 1858, with ''Sarcopyrenia gibba'' assigned as the type species. Sarcopyreniaceae is one of the few families composed entirely of lichenicolous fungi. Species Host data is from Diederich, Lawrey, and Ertz's 2018 review of lichenicolous fungi. *''Sarcopyrenia acutispora'' – on '' Bagliettoa calciseda'' *''Sarcopyrenia bacillosa'' – on saxicolous lichens *''Sarcopyrenia bacillospora'' – on '' Thelidium'' and '' Verrucaria'' *''Sarcopyrenia baetica'' – on ''Lecania'' cf. ''erysibe'' *''Sarcopyrenia beckhausiana'' – on saxicolous lichens (mostly from family Verrucariaceae) *''Sarcopyrenia calcarea'' *''Sarcopyrenia cylindrospora'' – mostly on '' Aspicilia'' *''Sarcopyrenia gibba'' – on saxicolous lichens *''Sarcopyrenia ...
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Kalbiana
''Kalbiana'' is a genus of saxicolous (rock-dwelling), crustose lichen of uncertain familial placement in the order Verrucariales. Both the genus and its only species, ''Kalbiana brasiliensis'', were described in 1988 by lichenologist Aino Henssen. The genus name honours German lichenologist Klaus Kalb Klaus Kalb (born 1942) is a German lichenologist and an authority on tropical lichens. Biography Klaus Kalb was born in Nuremberg in 1942 and grew up in southern Bavaria. From 1960 to 1965 he studied biology, chemistry, and geography at t ..., while the species epithet refers to Brazil, where the lichen is found. References Verrucariales Lichen genera Taxa described in 1988 Monotypic Eurotiomycetes genera Taxa named by Aino Henssen {{Eurotiomycetes-stub ...
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Gemmaspora
''Gemmaspora'' is a single-species fungal genus of uncertain familial placement in the order Verrucariales. It contains ''Gemmaspora lecanorae'', a lichenicolous (lichen A lichen ( , ) is a composite organism that arises from algae or cyanobacteria living among filaments of multiple fungi species in a mutualistic relationship.parasitises the lichen genus '' Aspicilia''. The genus was proposed in 2007 by David Hawksworth and Gökhan Halici to contain the fungus formerly known as ''Adelococcus lecanorae''. This species, originally
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Verrucaria
''Verrucaria'' is a genus of lichenized (lichen-forming) fungi in the family Verrucariaceae. Taxonomy The genus was circumscribed by German botanist Heinrich Adolph Schrader in 1794, with '' Verrucaria rupestris'' assigned as the type species. In his brief diagnosis of the genus, Schrader mentioned the more or less spherical (''subglobose''), closed ascomata, and the crustose thallus. The genus name is derived from the Latin word ''verruca'' (meaning "wart") and the suffix ''-aria'' (meaning "belonging to" or "possession"). Ecology As of 2015, there were 16 ''Verrucaria'' species classified as marine species: '' V. adguttata'', '' V. allantoidea'', '' V. ceuthocarpa'', '' V. corallensi'', '' V. ditmarsica'', '' V. erichsenii'', '' V. halizoa'', '' V. halochlora'', '' V. microsporoides'', '' V. paulula'', '' V. psychrophila'', '' V. sandstedei'', '' V. serpuloides'', '' V. sessilis'', '' V. subdiscreta'', ...
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Lichen
A lichen ( , ) is a composite organism that arises from algae or cyanobacteria living among filaments of multiple fungi species in a mutualistic relationship.Introduction to Lichens – An Alliance between Kingdoms
. University of California Museum of Paleontology.
Lichens have properties different from those of their component organisms. They come in many colors, sizes, and forms and are sometimes plant-like, but are not s. They may have tiny, leafless branches (); flat leaf-like structures (

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Lichen Orders
A lichen ( , ) is a composite organism that arises from algae or cyanobacteria living among filaments of multiple fungi species in a mutualistic relationship.Introduction to Lichens – An Alliance between Kingdoms
. University of California Museum of Paleontology.
Lichens have properties different from those of their component organisms. They come in many colors, sizes, and forms and are sometimes plant-like, but are not s. They may have tiny, leafless branches (); flat leaf-like structures (