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Tremble Dance
A tremble dance is a dance performed by forager honey bees of the species ''Apis mellifera'' to recruit more receiver honey bees to collect nectar from the workers. History of discovery The tremble dance was first described by Karl von Frisch in the 1920s (who was also first to describe the waggle dance), but no light was shed on its function until 1993 when Wolfgang Kirschner discovered that, when performed, the dance stopped nearby workers from flying to gather more nectar. Function The tremble dance of the honeybee is similar to the waggle dance, but is used by a forager when the foraging bee perceives a long delay in unloading its nectar or a shortage of receiver bees, sometimes due to low numbers of receiver bees. It may also spread the scent released during the forager's waggle dance. Like the waggle dance, the tremble dance is likely one of two "primary regulation mechanisms" for regulating bee colony behavior at the group level, and one of four or five observed mechanisms ...
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Honey Bee
A honey bee (also spelled honeybee) is a eusocial flying insect within the genus ''Apis'' of the bee clade, all native to Afro-Eurasia. After bees spread naturally throughout Africa and Eurasia, humans became responsible for the current cosmopolitan distribution of honey bees, introducing multiple subspecies into South America (early 16th century), North America (early 17th century), and Australia (early 19th century). Honey bees are known for their construction of perennial colonial nests from wax, the large size of their colonies, and surplus production and storage of honey, distinguishing their hives as a prized foraging target of many animals, including honey badgers, bears and human hunter-gatherers. Only eight surviving species of honey bee are recognized, with a total of 43 subspecies, though historically 7 to 11 species are recognized. Honey bees represent only a small fraction of the roughly 20,000 known species of bees. The best known honey bee is the weste ...
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Western Honey Bee
The western honey bee or European honey bee (''Apis mellifera'') is the most common of the 7–12 species of honey bees worldwide. The genus name ''Apis'' is Latin for "bee", and ''mellifera'' is the Latin for "honey-bearing" or "honey carrying", referring to the species' production of honey. Like all honey bee species, the western honey bee is eusocial, creating colonies with a single fertile female (or "queen"), many normally non-reproductive females or "workers", and a small proportion of fertile males or " drones". Individual colonies can house tens of thousands of bees. Colony activities are organized by complex communication between individuals, through both pheromones and the dance language. The western honey bee was one of the first domesticated insects, and it is the primary species maintained by beekeepers to this day for both its honey production and pollination activities. With human assistance, the western honey bee now occupies every continent except Antarctica. W ...
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Nectar
Nectar is a sugar-rich liquid produced by plants in glands called nectaries or nectarines, either within the flowers with which it attracts pollinating animals, or by extrafloral nectaries, which provide a nutrient source to animal mutualists, which in turn provide herbivore protection. Common nectar-consuming pollinators include mosquitoes, hoverflies, wasps, bees, butterflies and moths, hummingbirds, honeyeaters and bats. Nectar plays a crucial role in the foraging economics and evolution of nectar-eating species; for example, nectar foraging behavior is largely responsible for the divergent evolution of the African honey bee, ''A. m. scutellata'' and the western honey bee. Nectar is an economically important substance as it is the sugar source for honey. It is also useful in agriculture and horticulture because the adult stages of some predatory insects feed on nectar. For example, a number of parasitoid wasps (e.g. the social wasp species ''Apoica flavissima'') rely ...
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Worker Bee
A worker bee is any female (eusocial) bee that lacks the full reproductive capacity of the colony's queen bee; under most circumstances, this is correlated to an increase in certain non-reproductive activities relative to a queen. While worker bees occur in all eusocial bee species, the term is rarely used (outside of scientific literature) for any bees other than honey bees. Honey bee workers gather pollen into the pollen baskets on their back legs and carry it back to the hive where it is used as food for the developing brood. Pollen carried on their bodies may be carried to another flower where a small portion can rub off onto the pistil, resulting in cross pollination. A significant amount of the world's food supply, particularly fruit, depends greatly on crop pollination by honey bees. Nectar is sucked up through the proboscis, mixed with enzymes in the stomach, and carried back to the hive, where it is stored in wax cells and evaporated into honey. Life cycle Although the ...
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Karl Von Frisch
Karl Ritter von Frisch, (20 November 1886 – 12 June 1982) was a German-Austrian ethologist who received the Nobel Prize in Physiology or Medicine in 1973, along with Nikolaas Tinbergen and Konrad Lorenz. His work centered on investigations of the sensory perceptions of the honey bee and he was one of the first to translate the meaning of the waggle dance. His theory, described in his 1927 book ''Aus dem Leben der Bienen'' (translated into English as ''The Dancing Bees''), was disputed by other scientists and greeted with skepticism at the time. Only much later was it shown to be an accurate theoretical analysis. Early life Karl von Frisch was the son of the surgeon and urologist Anton von Frisch (1849–1917), by his marriage to Marie Exner. Karl was the youngest of four sons, all of whom became university professors. Von Frisch was of partial Jewish heritage. Karl studied in Vienna under Hans Leo Przibram and in Munich under Richard von Hertwig, initially in the field of med ...
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Waggle Dance
Waggle dance is a term used in beekeeping and ethology for a particular figure-eight dance of the honey bee. By performing this dance, successful foragers can share information about the direction and distance to patches of flowers yielding nectar and pollen, to water sources, or to new nest-site locations with other members of the colony. The waggle dance and the round dance are two forms of dance behaviour that are part of a continuous transition. As the distance between the resource and the hive increases, the round dance transforms into variations of a transitional dance, which, when communicating resources at even greater distances, becomes the waggle dance. In the case of ''Apis mellifera ligustica'', the round dance is performed until the resource is about 10 metres away from the hive, transitional dances are performed when the resource is at a distance of 20 to 30 metres away from the hive, and finally, when it is located at distances greater than 40 metres from the hive ...
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Wolfgang Kirschner
Wolfgang is a German male given name traditionally popular in Germany, Austria and Switzerland. The name is a combination of the Old High German words ''wolf'', meaning "wolf", and '' gang'', meaning "path", "journey", "travel". Besides the regular "wolf", the first element also occurs in Old High German as the combining form "-olf". The earliest reference of the name being used was in the 8th century. The name was also attested as "Vulfgang" in the Reichenauer Verbrüderungsbuch in the 9th century. The earliest recorded famous bearer of the name was a tenth-century Saint Wolfgang of Regensburg. Due to the lack of conflict with the pagan reference in the name with Catholicism, it is likely a much more ancient name whose meaning had already been lost by the tenth century. Grimm ('' Teutonic Mythology'' p. 1093) interpreted the name as that of a hero in front of whom walks the "wolf of victory". A Latin gloss by Arnold of St Emmeram interprets the name as ''Lupambulus''.E. Fö ...
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Ethanol
Ethanol (abbr. EtOH; also called ethyl alcohol, grain alcohol, drinking alcohol, or simply alcohol) is an organic compound. It is an Alcohol (chemistry), alcohol with the chemical formula . Its formula can be also written as or (an ethyl group linked to a hydroxyl group). Ethanol is a Volatility (chemistry), volatile, Combustibility and flammability, flammable, colorless liquid with a characteristic wine-like odor and pungent taste. It is a psychoactive recreational drug, the active ingredient in alcoholic drinks. Ethanol is naturally produced by the fermentation process of Carbohydrate, sugars by yeasts or via Petrochemistry, petrochemical processes such as ethylene hydration. It has medical applications as an antiseptic and disinfectant. It is used as a chemical solvent and in the Chemical synthesis, synthesis of organic compounds, and as a Alcohol fuel, fuel source. Ethanol also can be dehydrated to make ethylene, an important chemical feedstock. As of 2006, world produ ...
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Bee Learning And Communication
Bee learning and communication includes cognitive and sensory processes in all kinds of bees, that is the insects in the seven families making up the clade Anthophila. Some species have been studied more extensively than others, in particular ''Apis mellifera'', or European honey bee. Color learning has also been studied in bumblebees. Honey bees are sensitive to odors (including pheromones), tastes, and colors, including ultraviolet. They can demonstrate capabilities such as color discrimination through classical and operant conditioning and retain this information for several days at least; they communicate the location and nature of sources of food; they adjust their foraging to the times at which food is available; they may even form cognitive maps of their surroundings. They also communicate with each other by means of a "waggle dance" and in other ways. Learning Honey bees are adept at associative learning, and many of the phenomena of operant and classical conditioning ...
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Queen Bee
A queen bee is typically an adult, mated female (gyne) that lives in a colony or hive of honey bees. With fully developed reproductive organs, the queen is usually the mother of most, if not all, of the bees in the beehive. Queens are developed from larvae selected by worker bees and specially fed in order to become sexually mature. There is normally only one adult, mated queen in a hive, in which case the bees will usually follow and fiercely protect her. The term "queen bee" can be more generally applied to any dominant reproductive female in a colony of a eusocial bee species other than honey bees. However, as in the Brazilian stingless bee ''Schwarziana quadripunctata'', a single nest may have multiple queens or even dwarf queens, ready to replace a dominant queen in a case of sudden death. Development During the warm parts of the year, female "worker" bees leave the hive every day to collect nectar and pollen. While male bees serve no architectural or pollinating purpo ...
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Eusociality
Eusociality (from Greek εὖ ''eu'' "good" and social), the highest level of organization of sociality, is defined by the following characteristics: cooperative brood care (including care of offspring from other individuals), overlapping generations within a colony of adults, and a division of labor into reproductive and non-reproductive groups. The division of labor creates specialized behavioral groups within an animal society which are sometimes referred to as 'castes'. Eusociality is distinguished from all other social systems because individuals of at least one caste usually lose the ability to perform at least one behavior characteristic of individuals in another caste. Eusocial colonies can be viewed as superorganisms. Eusociality exists in certain insects, crustaceans, and mammals. It is mostly observed and studied in the Hymenoptera (ants, bees, and wasps) and in Blattodea (termites). A colony has caste differences: queens and reproductive males take the roles of the ...
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Grooming Dance
A grooming dance, grooming invitation dance or shaking dance is a dance performed by honeybees to initiate social grooming, allogrooming. It was first reported in 1945 by biologist Mykola H. Hadak. An increase in the frequency of the grooming dance has been observed among the bees of mite-infested colonies, and among bees who have been dusted with small particles of chalk dust.Land, B. B., T. D. Seeley. (28 January 2004) The Grooming Invitation Dance of the Honey Bee. ''Ethology.'' Volume 110, Issue 1. pp. 1-10. See also *Bee learning and communication *Tremble dance *Waggle dance References

{{Neuroethology Animal communication Western honey bee behavior Neuroethology ...
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