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Theriognathus
''Theriognathus'' (from Greek therion: beast, mammal, Greek, gnathos, “jaw,” +us, pronounced THEH-ree-OG-nah-thuss) is an extinct genus of therocephalian therapsid belonging to the family Whaitsiidae, known from fossils from South Africa, Zambia, and Tanzania. ''Theriognathus'' has been dated as existing during the Late Permian. Although ''Theriognathus'' means mammal jaw, the lower jaw is actually made up of several bones as seen in modern reptiles, in contrast to mammals. ''Theriognathus'' displayed many different reptilian and mammalian characteristics. For example, ''Theriognathus'' had canine teeth like mammals, and a secondary palate, multiple bones in the mandible, and a typical reptilian jaw joint, all characteristics of reptiles. It is speculated that ''Theriognathus'' was either carnivorous or omnivorous based on its teeth, and was suited to hunting small prey in undergrowth. This synapsid adopted a sleek profile of a mammalian predator, with a narrow snout and aro ...
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Notosollasia
''Theriognathus'' (from Greek therion: beast, mammal, Greek, gnathos, “jaw,” +us, pronounced THEH-ree-OG-nah-thuss) is an extinct genus of therocephalian therapsid belonging to the family Whaitsiidae, known from fossils from South Africa, Zambia, and Tanzania. ''Theriognathus'' has been dated as existing during the Late Permian. Although ''Theriognathus'' means mammal jaw, the lower jaw is actually made up of several bones as seen in modern reptiles, in contrast to mammals. ''Theriognathus'' displayed many different reptilian and mammalian characteristics. For example, ''Theriognathus'' had canine teeth like mammals, and a secondary palate, multiple bones in the mandible, and a typical reptilian jaw joint, all characteristics of reptiles. It is speculated that ''Theriognathus'' was either carnivorous or omnivorous based on its teeth, and was suited to hunting small prey in undergrowth. This synapsid adopted a sleek profile of a mammalian predator, with a narrow snout and aro ...
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Theriognathus
''Theriognathus'' (from Greek therion: beast, mammal, Greek, gnathos, “jaw,” +us, pronounced THEH-ree-OG-nah-thuss) is an extinct genus of therocephalian therapsid belonging to the family Whaitsiidae, known from fossils from South Africa, Zambia, and Tanzania. ''Theriognathus'' has been dated as existing during the Late Permian. Although ''Theriognathus'' means mammal jaw, the lower jaw is actually made up of several bones as seen in modern reptiles, in contrast to mammals. ''Theriognathus'' displayed many different reptilian and mammalian characteristics. For example, ''Theriognathus'' had canine teeth like mammals, and a secondary palate, multiple bones in the mandible, and a typical reptilian jaw joint, all characteristics of reptiles. It is speculated that ''Theriognathus'' was either carnivorous or omnivorous based on its teeth, and was suited to hunting small prey in undergrowth. This synapsid adopted a sleek profile of a mammalian predator, with a narrow snout and aro ...
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Whaitsiidae
Whaitsiidae is an extinct family of therocephalian therapsid Therapsida is a major group of eupelycosaurian synapsids that includes mammals, their ancestors and relatives. Many of the traits today seen as unique to mammals had their origin within early therapsids, including limbs that were oriented more ...s. References Permian first appearances Permian extinctions Prehistoric therapsid families {{paleo-therapsid-stub ...
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Therocephalian
Therocephalia is an extinct suborder of eutheriodont therapsids (mammals and their close relatives) from the Permian and Triassic. The therocephalians ("beast-heads") are named after their large skulls, which, along with the structure of their teeth, suggest that they were carnivores. Like other non-mammalian synapsids, therocephalians were once described as "mammal-like reptiles". Therocephalia is the group most closely related to the cynodonts, which gave rise to the mammals. This relationship takes evidence in a variety of skeletal features. The fossils of therocephalians are numerous in the Karoo of South Africa, but have also been found in Russia, China, Tanzania, Zambia, and Antarctica. Early therocephalian fossils discovered in Middle Permian deposits of South Africa support a Gondwanan origin for the group, which seems to have spread quickly across Earth. Although almost every therocephalian lineage ended during the great Permian–Triassic extinction event, a few represen ...
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Wuchiapingian
In the geologic timescale, the Wuchiapingian or Wujiapingian (from in the Liangshan area of Hanzhong, Shaanxi Province) is an age or stage of the Permian. It is also the lower or earlier of two subdivisions of the Lopingian Epoch or Series. The Wuchiapingian spans the time between and million years ago (Ma). It was preceded by the Capitanian and followed by the Changhsingian. Regional stages with which the Wuchiapingian is coeval or overlaps include the Djulfian or Dzhulfian, Longtanian, Rustlerian, Saladoan, and Castilian. Stratigraphic definitions The Wuchiapingian was first used in 1962, when the Lopingian Series of southwestern China was divided in the Changhsingian and Wuchiapingian Formations. In 1973 the Wuchiapingian was first used as a chronostratigraphic unit (i.e. a stage, as opposed to a formation, which is a lithostratigraphic unit). The base of the Wuchiapingian Stage is defined as the place in the stratigraphic record where the conodont species ''Clarkina po ...
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Beaufort Group
The Beaufort Group is the third of the main subdivisions of the Karoo Supergroup in South Africa. It is composed of a lower Adelaide Subgroup and an upper Tarkastad Subgroup. It follows conformably after the Ecca Group and unconformably underlies the Stormberg Group. Based on stratigraphic position, lithostratigraphic and biostratigraphic correlations, palynological analyses, and other means of geological dating, the Beaufort Group rocks are considered to range between Middle Permian (Wordian) to Early Triassic (Anisian) in age. Background During the period when sedimentation of the Beaufort Group rocks took place, the Ecca sea had retreated to the northeastern Karoo Basin. All sediment deposition at this time took place in a terrestrial, although in a predominantly fluvial or alluvial environment that was seasonally arid. This environment covered a vast area and deposition was influenced by a retroarc foreland basin. This foreland system was caused by crustal uplift (oro ...
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Guadalupian
The Guadalupian is the second and middle series/epoch of the Permian. The Guadalupian was preceded by the Cisuralian and followed by the Lopingian. It is named after the Guadalupe Mountains of New Mexico and Texas, and dates between 272.95 ± 0.5 – 259.1 ± 0.4 Mya. The series saw the rise of the therapsids, a minor extinction event called Olson's Extinction and a significant mass extinction called the end-Capitanian extinction event. The Guadalupian was previously known as the Middle Permian. Name and background The Guadalupian is the second and middle series or epoch of the Permian. Previously called Middle Permian, the name of this epoch is part of a revision of Permian stratigraphy for standard global correlation. The name "Guadalupian" was first proposed in the early 1900s, and approved by the International Subcommission on Permian Stratigraphy in 1996. References to the Middle Permian still exist. The Guadalupian was preceded by the Cisuralian and followed by the Lop ...
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Cladistics
Cladistics (; ) is an approach to biological classification in which organisms are categorized in groups (" clades") based on hypotheses of most recent common ancestry. The evidence for hypothesized relationships is typically shared derived characteristics ( synapomorphies'')'' that are not present in more distant groups and ancestors. However, from an empirical perspective, common ancestors are inferences based on a cladistic hypothesis of relationships of taxa whose character states can be observed. Theoretically, a last common ancestor and all its descendants constitute a (minimal) clade. Importantly, all descendants stay in their overarching ancestral clade. For example, if the terms ''worms'' or ''fishes'' were used within a ''strict'' cladistic framework, these terms would include humans. Many of these terms are normally used paraphyletically, outside of cladistics, e.g. as a 'grade', which are fruitless to precisely delineate, especially when including extinct species. R ...
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Paraphyletic
In taxonomy (general), taxonomy, a group is paraphyletic if it consists of the group's most recent common ancestor, last common ancestor and most of its descendants, excluding a few Monophyly, monophyletic subgroups. The group is said to be paraphyletic ''with respect to'' the excluded subgroups. In contrast, a monophyletic group (a clade) includes a common ancestor and ''all'' of its descendants. The terms are commonly used in phylogenetics (a subfield of biology) and in the tree model of historical linguistics. Paraphyletic groups are identified by a combination of Synapomorphy and apomorphy, synapomorphies and symplesiomorphy, symplesiomorphies. If many subgroups are missing from the named group, it is said to be polyparaphyletic. The term was coined by Willi Hennig to apply to well-known taxa like Reptilia (reptiles) which, as commonly named and traditionally defined, is paraphyletic with respect to mammals and birds. Reptilia contains the last common ancestor of reptiles a ...
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Charassognathus
''Charassognathus'' (meaning 'notched jaw') is an extinct genus of Late Permian cynodonts. Described in 2007 from a locality near Fraserburg, South Africa, ''Charassognathus'' is one of the earliest and most basal cynodonts. It is known only from the holotype, which dates from the Late Permian Period. The type and only species is ''C. gracilis''. The holotype ( SAM-PK-K 10369), found in the ''Tropidostoma'' Assemblage Zone of the Teekloof Formation, is made up of a crushed skull, partial lower jaw and one leg. ''Charassognathus'' was a quadrupedal predator. It was named for a notch on its coronoid process which most likely was the insertion point for a chewing muscle, the adductor mandibulae externus. ''Charassognathus'' was a small animal, with a skull only in length. Since the body of ''Charassognathus'' has not been discovered, its full length remains unknown, but estimates have been made at . Description ''Charassognathus'' has a snout that makes up slightly less th ...
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Cynodont
The cynodonts () (clade Cynodontia) are a clade of eutheriodont therapsids that first appeared in the Late Permian (approximately 260 mya), and extensively diversified after the Permian–Triassic extinction event. Cynodonts had a wide variety of lifestyles, including carnivory and herbivory. Mammals are cynodonts, as are their extinct ancestors and close relatives, having evolved from advanced probainognathian cynodonts during the Late Triassic. All other cynodont lines went extinct, with the last known non-mammalian cynodont group, the Tritylodontidae, having its youngest records in the Early Cretaceous. Description Early cynodonts have many of the skeletal characteristics of mammals. The teeth were fully differentiated and the braincase bulged at the back of the head. Outside of some crown-group mammals (notably the therians), all cynodonts probably laid eggs. The temporal fenestrae were much larger than those of their ancestors, and the widening of the zygomatic arch in ...
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Dicynodont
Dicynodontia is an extinct clade of anomodonts, an extinct type of non-mammalian therapsid. Dicynodonts were herbivorous animals with a pair of tusks, hence their name, which means 'two dog tooth'. Members of the group possessed a horny, typically toothless beak, unique amongst all synapsids. Dicynodonts first appeared in Southern Pangaea during the mid-Permian, ca. 270–260 million years ago, and became globally distributed and the dominant herbivorous animals in the Late Permian, ca. 260–252 Mya. They were devastated by the end-Permian Extinction that wiped out most other therapsids ca. 252 Mya. They rebounded during the Triassic but died out towards the end of that period. They were the most successful and diverse of the non-mammalian therapsids, with over 70 genera known, varying from rat-sized burrowers to elephant-sized browsers. Characteristics The dicynodont skull is highly specialised, light but strong, with the synapsid temporal openings at the rear of the skull ...
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