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Guadalupian
The Guadalupian is the second and middle series/epoch of the Permian. The Guadalupian was preceded by the Cisuralian and followed by the Lopingian. It is named after the Guadalupe Mountains of New Mexico and Texas, and dates between 272.95 ± 0.5 – 259.1 ± 0.4 Mya. The series saw the rise of the therapsids, a minor extinction event called Olson's Extinction and a significant mass extinction called the end-Capitanian extinction event. The Guadalupian was previously known as the Middle Permian. Name and background The Guadalupian is the second and middle series or epoch of the Permian. Previously called Middle Permian, the name of this epoch is part of a revision of Permian stratigraphy for standard global correlation. The name "Guadalupian" was first proposed in the early 1900s, and approved by the International Subcommission on Permian Stratigraphy in 1996. References to the Middle Permian still exist. The Guadalupian was preceded by the Cisuralian and followed by the Lop ...
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End-Capitanian Extinction Event
The Capitanian mass extinction event, also known as the end-Guadalupian extinction event or the pre-Lopingian crisis was an extinction event that predated the end-Permian extinction event and occurred around 260 million years ago during a period of decreased species richness and increased extinction rates in the late Middle Permian during the Guadalupian epoch. It is often called the end-Guadalupian extinction event because of its initial recognition between the Guadalupian and Lopingian series; however, more refined stratigraphic study suggests that extinction peaks in many taxonomic groups occurred within the Guadalupian, in the latter half of the Capitanian age.Bond, D. P. G., Wignall, P. B., Wang, W., Izon, G., Jiang, H. S., Lai, X. L., Sund, Y.-D., Newtona, R.J., Shaoe, L.-Y., Védrinea, S. & Cope, H. (2010). "The mid-Capitanian (Middle Permian) mass extinction and carbon isotope record of South China". ''Palaeogeography, Palaeoclimatology, Palaeoecology'', 292 (1-2), pp. 282- ...
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Permian
The Permian ( ) is a geologic period and stratigraphic system which spans 47 million years from the end of the Carboniferous Period million years ago (Mya), to the beginning of the Triassic Period 251.9 Mya. It is the last period of the Paleozoic Era; the following Triassic Period belongs to the Mesozoic Era. The concept of the Permian was introduced in 1841 by geologist Sir Roderick Murchison, who named it after the region of Perm in Russia. The Permian witnessed the diversification of the two groups of amniotes, the synapsids and the sauropsids ( reptiles). The world at the time was dominated by the supercontinent Pangaea, which had formed due to the collision of Euramerica and Gondwana during the Carboniferous. Pangaea was surrounded by the superocean Panthalassa. The Carboniferous rainforest collapse left behind vast regions of desert within the continental interior. Amniotes, which could better cope with these drier conditions, rose to dominance in place of their am ...
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Permian–Triassic Extinction Event
The Permian–Triassic (P–T, P–Tr) extinction event, also known as the Latest Permian extinction event, the End-Permian Extinction and colloquially as the Great Dying, formed the boundary between the Permian and Triassic geologic periods, as well as between the Paleozoic and Mesozoic eras, approximately 251.9 million years ago. It is the Earth's most severe known extinction event, with the extinction of 57% of biological families, 83% of genera, 81% of marine species and 70% of terrestrial vertebrate species. It was the largest known mass extinction of insects. There is evidence for one to three distinct pulses, or phases, of extinction. The scientific consensus is that the main cause of extinction was the large amount of carbon dioxide emitted by the volcanic eruptions that created the Siberian Traps, which elevated global temperatures, and in the oceans led to widespread anoxia and acidification. Proposed contributing factors include: the emission of much addition ...
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Olson's Extinction
Olson's Extinction was a mass extinction that occurred in the late Cisuralian or early Guadalupian of the Permian period and which predated the Permian–Triassic extinction event. It is named after Everett C. Olson. There was a sudden change between the early Permian and middle/late Permian faunas. Some authors also place a hiatus in the continental fossil record around that time, but others disagree. Since then this event has been realized across many groups, including plants, marine invertebrates, and tetrapods. Identification The first evidence of extinction came when Everett C. Olson noted a hiatus between early Permian faunas dominated by pelycosaurs and therapsid-dominated faunas of the middle and late Permian. First considered to be a preservational gap in the fossil record, the event was originally dubbed 'Olson's Gap'. To compound the difficulty in identifying the cause of the 'gap', researchers were having difficulty in resolving the uncertainty which exists regardi ...
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Jinogondolella
''Jinogondolella'' is an extinct genus of conodonts.Mesogondolella and Jinogondolella (Conodonta): Multielement definition of the taxa that bracket the basal Guadalupian (Middle Permian Series) GSSP. Lance L. Lambert, Bruce R. Wardlaw and Charles M. Henderson, Palaeoworld, Volume 16, Issues 1–3, January–September 2007, Pages 208–221, Use in stratigraphy The top of the Kungurian (the base of the Roadian and the Guadalupian series) is defined as the place in the stratigraphic record where fossils of the species ''Jinogondolella nanginkensis'' first appeared. The base of the Capitanian stage is defined as the place in the stratigraphic record where fossils of the species ''Jinogondolella postserrata'' first appeared. The global reference profile for this stratigraphic boundary is located at Nipple Hill in the southern Guadalupe Mountains of Texas. The base of the Wordian In the geologic timescale, the Wordian is an age or stage of the Permian. It is the middle of th ...
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Cisuralian
The Cisuralian is the first Series (stratigraphy), series/Epoch (geology), epoch of the Permian. The Cisuralian was preceded by the Pennsylvanian (geology), Pennsylvanian and followed by the Guadalupian. The Cisuralian Epoch is named after the western slopes of the Ural Mountains in Russia and Kazakhstan and dates between 298.9 ± 0.15 – 272.3 ± 0.5 Mya. The Cisuralian is often synonymous with the informal terms early Permian or lower Permian. It corresponds approximately with the Wolfcampian in southwestern North America. The series saw the appearance of beetles and Fly, flies and was a relatively stable warming period of about 21 million years. Name and background The Cisuralian is the first series or epoch of the Permian. The Cisuralian was preceded by the last Pennsylvanian epoch (Gzhelian) and is followed by the Permian Guadalupian Epoch. The name "Cisuralian" was proposed in 1982, and approved by the International Subcommission on Permian Stratigraphy in 1996. The Ci ...
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Mass Extinction
An extinction event (also known as a mass extinction or biotic crisis) is a widespread and rapid decrease in the biodiversity on Earth. Such an event is identified by a sharp change in the diversity and abundance of multicellular organisms. It occurs when the rate of extinction increases with respect to the background extinction rate and the rate of speciation. Estimates of the number of major mass extinctions in the last 540 million years range from as few as five to more than twenty. These differences stem from disagreement as to what constitutes a "major" extinction event, and the data chosen to measure past diversity. The "Big Five" mass extinctions In a landmark paper published in 1982, Jack Sepkoski and David M. Raup identified five particular geological intervals with excessive diversity loss. They were originally identified as outliers on a general trend of decreasing extinction rates during the Phanerozoic, but as more stringent statistical tests have been applied to ...
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Theriodont
The theriodonts (clade Theriodontia) are a major group of therapsids which appeared during the Middle Permian and which includes the gorgonopsians and the eutheriodonts, itself including the therocephalians and the cynodonts. Naming In 1876, Richard Owen named a suborder Theriodontia, which he divided into the Cynodontia and the Gomphodontia. The modern clade concept was devised by James Allen Hopson. In his system, Theriodontia fall into two main groups: the Gorgonopsia and the Eutheriodontia. The latter consist of the Therocephalia and Cynodontia.J.A. Hopson. 1999. "Therapsids". ''Encyclopedia of Paleontology'' 2: 1256-1266 Evolution Theriodonts appeared at the same time as their sister group within the Neotherapsida, the Anomodontia, about 270 million years ago, in the Middle Permian. Even these early theriodonts were more mammal-like than their anomodont and dinocephalian contemporaries. Early theriodonts may have been warm-blooded. Early forms were carnivorous, but se ...
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Anomodont
Anomodontia is an extinct group of non-mammalian therapsids from the Permian and Triassic periods. By far the most speciose group are the dicynodonts, a clade of beaked, tusked herbivores.Chinsamy-Turan, A. (2011) ''Forerunners of Mammals: Radiation - Histology - Biology''p.39.Indiana University Press, . Retrieved May 2012 Anomodonts were very diverse during the Middle Permian, including primitive forms like ''Anomocephalus'' and ''Patranomodon'' and groups like Venyukovioidea and Dromasauria. Dicynodonts became the most successful and abundant of all herbivores in the Late Permian, filling ecological niches ranging from large browsers down to small burrowers. Few dicynodont families survived the Permian–Triassic extinction event, but one lineage (Kannemeyeriiformes) evolved into large, stocky forms that became dominant terrestrial herbivores right until the Late Triassic, when changing conditions caused them to decline, finally going extinct during the Triassic-Jurassic ext ...
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Epoch (geology)
The geologic time scale, or geological time scale, (GTS) is a representation of time based on the rock record of Earth. It is a system of chronological dating that uses chronostratigraphy (the process of relating strata to time) and geochronology (scientific branch of geology that aims to determine the age of rocks). It is used primarily by Earth scientists (including geologists, paleontologists, geophysicists, geochemists, and paleoclimatologists) to describe the timing and relationships of events in geologic history. The time scale has been developed through the study of rock layers and the observation of their relationships and identifying features such as lithologies, paleomagnetic properties, and fossils. The definition of standardized international units of geologic time is the responsibility of the International Commission on Stratigraphy (ICS), a constituent body of the International Union of Geological Sciences (IUGS), whose primary objective is to precisely define gl ...
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Guadalupe Mountains
The Guadalupe Mountains ( es, Sierra de Guadalupe) are a mountain range located in West Texas and southeastern New Mexico. The range includes the highest summit in Texas, Guadalupe Peak, , and the "signature peak" of West Texas, El Capitan, both of which are located within Guadalupe Mountains National Park. The Guadalupe Mountains are bordered by the Pecos River valley and Llano Estacado to the east and north, Delaware Mountains to the south, and Sacramento Mountains to the west. One of the clearest exposures of a prehistoric reef is preserved in the mountain range's bedrock geology. Bedrock contains fossils of reef-dwelling organisms from the Permian period, and the geology is widely studied, mostly by stratigraphers, paleontologists, and Paleoecologists (see geology section). History Archaeological evidence has shown that people lived over 10,000 years ago in and among the many caves and alcoves. The first humans to live here were hunter-gatherers who followed large game ...
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Dinocephalia
Dinocephalians (terrible heads) are a clade of large-bodied early therapsids that flourished in the Early and Middle Permian between 279.5 and 260 million years ago (Ma), but became extinct during the Capitanian mass extinction event. Dinocephalians included herbivorous, carnivorous, and omnivorous forms. Many species had thickened skulls with many knobs and bony projections. Dinocephalians were the first non-mammalian therapsids to be scientifically described and their fossils are known from Russia, China, Brazil, South Africa, Zimbabwe, and Tanzania. Description Apart from the Biarmosuchians, the dinocephalians are the least advanced therapsids, although still uniquely specialised in their own way. They retain a number of primitive characteristics (e.g. no secondary palate, small dentary) shared with their pelycosaur ancestors, although they are also more advanced in possessing therapsid adaptations like the expansion of the ilium and more erect limbs. They include carni ...
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