Stockmansella
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Stockmansella
''Stockmansella'' is a genus of extinct plants of the Middle Devonian (Eifelian stage, around ), fossils of which have been found in north-west Germany. The sporophyte generation consists of prostrate dichotomizing stems (axes) up to 10 cm long and around 3mm wide, which at intervals produce narrower smooth upright stems. These bear sporangia (spore-forming organs) on short lateral branches (sporangiophores). The prostrate stems have bulges from which rhizoids form. Both prostrate and upright stems have a central strand of conducting tissue which contains simple tracheids, so that ''Stockmansella'' is a vascular plant. The genus was created by Fairon-Demaret for fossil forms previously assigned to ''Taeniocrada'' but which differ in having single lateral sporangia. (She initially gave the genus the name ''Stockmansia'', but this had already been used for a genus of ferns.) The form genus ''Sciadophyton'' is thought to be the gametophyte stage of several early land plants, i ...
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Rhyniopsida
The rhyniophytes are a group of extinct early vascular plants that are considered to be similar to the genus ''Rhynia'', found in the Early Devonian (around ). Sources vary in the name and rank used for this group, some treating it as the class Rhyniopsida, others as the subdivision Rhyniophytina or the division Rhyniophyta. The first definition of the group, under the name Rhyniophytina, was by Banks, since when there have been many redefinitions, including by Banks himself. "As a result, the Rhyniophytina have slowly dissolved into a heterogeneous collection of plants ... the group contains only one species on which all authors agree: the type species ''Rhynia gwynne-vaughanii''". When defined very broadly, the group consists of plants with dichotomously branched, naked aerial axes ("stems") with terminal spore-bearing structures (sporangia). The rhyniophytes are considered to be stem group tracheophytes (vascular plants). Definitions The group was described as a subdivision of ...
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Taeniocrada
''Taeniocrada'' is a genus of extinct plants of Devonian age (). It is used as a form genus for fossil plants with leafless flattened stems which divided dichotomously and had prominent midribs regarded as containing vascular tissues. It has been suggested that some species assigned to this genus were aquatic. Description Key features of the original definition of the genus ''Taeniocrada'' were that it possessed leafless flattened stems with prominent midribs which appeared to contain vascular tissues. As more species were added to the genus, its definition became less clear. Three of the better-known species are ''T. decheniana'', ''T. dubia'', and ''T. stilesvillensis''. ''Taeniocrada decheniana'', from the Lower Devonian, had separate fertile stems which repeatedly branched in a dichotomous fashion ending in sporangia between 3 and 7 mm long. A few sporangia were borne on the sides of stems. The species was found in dense stands. It has been suggested that it was aq ...
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Polysporangiophyte
Polysporangiophytes, also called polysporangiates or formally Polysporangiophyta, are plants in which the spore-bearing generation (sporophyte) has branching stems (axes) that bear sporangia. The name literally means 'many sporangia plant'. The clade includes all land plants (embryophytes) except for the bryophytes (liverworts, mosses and hornworts) whose sporophytes are normally unbranched, even if a few exceptional cases occur. While the definition is independent of the presence of vascular tissue, all living polysporangiophytes also have vascular tissue, i.e., are vascular plants or tracheophytes. Extinct polysporangiophytes are known that have no vascular tissue and so are not tracheophytes. Early polysporangiophytes History of discovery Paleobotanists distinguish between micro- and megafossils. Microfossils are primarily spores, either single or in groups. Megafossils are preserved parts of plants large enough to show structure, such as stem cross-sections or branching patte ...
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Rhynia
''Rhynia'' is a single-species genus of Devonian vascular plants. ''Rhynia gwynne-vaughanii'' was the sporophyte generation of a vascular, axial, free-sporing Alternation of generations, diplohaplontic embryophyte, embryophytic land plant of the Early Devonian that had plant anatomy, anatomical features more advanced than those of the bryophytes. ''Rhynia gwynne-vaughanii'' was a member of a sister group to all other tracheophyte, eutracheophytes, including modern vascular plants. Description ''Rhynia gwynne-vaughanii'' was first described as a new species by Robert Kidston and William Henry Lang, William H. Lang in 1917. The species is known only from the Rhynie chert in Aberdeenshire, Scotland, where it grew in the vicinity of a silica-rich hot spring. ''Rhynia'' was a vascular plant, and grew in association with other vascular plants such as ''Asteroxylon, Asteroxylon mackei'', a probable ancestor of modern clubmosses (Lycopsida), and with pre-vascular plants such as ''Aglao ...
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Species
In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. Other ways of defining species include their karyotype, DNA sequence, morphology, behaviour or ecological niche. In addition, paleontologists use the concept of the chronospecies since fossil reproduction cannot be examined. The most recent rigorous estimate for the total number of species of eukaryotes is between 8 and 8.7 million. However, only about 14% of these had been described by 2011. All species (except viruses) are given a two-part name, a "binomial". The first part of a binomial is the genus to which the species belongs. The second part is called the specific name or the specific epithet (in botanical nomenclature, also sometimes i ...
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Euphyllophytina
The euphyllophytes are a clade of plants within the tracheophytes (the vascular plants). The group may be treated as an unranked clade, a division under the name Euphyllophyta or a subdivision under the name Euphyllophytina. The euphyllophytes are characterized by the possession of true leaves ("megaphylls"), and comprise one of two major lineages of extant vascular plants. As shown in the cladogram below, the euphyllophytes have a sister relationship to the lycopodiophytes or lycopsids. Unlike the lycopodiophytes, which consist of relatively few presently living or extant taxa, the euphyllophytes comprise the vast majority of vascular plant lineages that have evolved since both groups shared a common ancestor more than 400 million years ago. The euphyllophytes consist of two lineages, the spermatophytes A spermatophyte (; ), also known as phanerogam (taxon Phanerogamae) or phaenogam (taxon Phaenogamae), is any plant that produces seeds, hence the alternative name seed plant. Sp ...
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Lycopodiophyta
The lycophytes, when broadly circumscribed, are a vascular plant (tracheophyte) subgroup of the kingdom Plantae. They are sometimes placed in a division Lycopodiophyta or Lycophyta or in a subdivision Lycopodiophytina. They are one of the oldest lineages of extant (living) vascular plants; the group contains extinct plants that have been dated from the Silurian (ca. 425 million years ago). Lycophytes were some of the dominating plant species of the Carboniferous period, and included tree-like species, although extant lycophytes are relatively small plants. The scientific names and the informal English names used for this group of plants are ambiguous. For example, "Lycopodiophyta" and the shorter "Lycophyta" as well as the informal "lycophyte" may be used to include the extinct zosterophylls or to exclude them. Description Lycophytes reproduce by spores and have alternation of generations in which (like other vascular plants) the sporophyte generation is dominant. Some lycophyte ...
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Huvenia
''Huvenia'' is a genus of extinct plants of the Early Devonian (Pragian or Siegenian stage, around ), found in slate deposits of the Rhenish Massif. The sporophyte generation consisted of leafless stems (axes), which appear to be flattened, and which branch dichotomously. The strand of conducting tissue contains simple tracheids, making this a vascular plant Vascular plants (), also called tracheophytes () or collectively Tracheophyta (), form a large group of land plants ( accepted known species) that have lignified tissues (the xylem) for conducting water and minerals throughout the plant. They al ... (tracheophyte). The sporangia (spore-forming organs) are borne on the ends of short branching stems (sporangiophores) rather than terminating main stems as in some other early land plants. Sporangia appear to be twisted, but it is not clear whether this feature was present in life or developed after death. In 2004, Crane et al. published a cladogram for the polysporangiop ...
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Tracheophyta
Vascular plants (), also called tracheophytes () or collectively Tracheophyta (), form a large group of land plants ( accepted known species) that have lignified tissues (the xylem) for conducting water and minerals throughout the plant. They also have a specialized non-lignified tissue (the phloem) to conduct products of photosynthesis. Vascular plants include the clubmosses, horsetails, ferns, gymnosperms (including conifers), and angiosperms (flowering plants). Scientific names for the group include Tracheophyta, Tracheobionta and Equisetopsida ''sensu lato''. Some early land plants (the rhyniophytes) had less developed vascular tissue; the term eutracheophyte has been used for all other vascular plants, including all living ones. Historically, vascular plants were known as "higher plants", as it was believed that they were further evolved than other plants due to being more complex organisms. However, this is an antiquated remnant of the obsolete scala naturae, and the term ...
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Aglaophyton
''Aglaophyton major'' (or more correctly ''Aglaophyton majus'') was the sporophyte generation of a diplohaplontic, pre-vascular, axial, free-sporing land plant of the Lower Devonian (Pragian stage, around ). It had anatomical features intermediate between those of the bryophytes and vascular plants or tracheophytes. ''A. major'' was first described by Kidston and Lang in 1920 as the new species ''Rhynia major''. The species is known only from the Rhynie chert in Aberdeenshire, Scotland, where it grew in the vicinity of a silica-rich hot spring, together with a number of associated vascular plants such as a smaller species ''Rhynia gwynne-vaughanii'' which may be interpreted as a representative of the ancestors of modern vascular plants and '' Asteroxylon mackei'', which was an ancestor of modern clubmosses (Lycopsida). Description The stems of ''Aglaophyton'' were round in cross-section, smooth, unornamented, and up to about 6mm in diameter. Kidston and Lang interpreted the pl ...
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Horneophytopsida
The Horneophytopsida, informally called horneophytes, are a class of extinct plants which consisted of branched stems without leaves, true roots or vascular tissue, found from the Late Silurian to the Early Devonian (around ). They are the simplest known polysporangiophytes, i.e. plants with sporophytes bearing many spore-forming organs (sporangia) on branched stems. They were formerly classified among the rhyniophytes, but it was later found that some of the original members of the group had simple vascular tissue and others did not. The group has also been treated as the division Horneophyta. In 2004, Crane et al. published a cladogram for the polysporangiophytes in which the Horneophytopsida are shown as the sister group of all other polysporangiophytes. One other former rhyniophyte, ''Aglaophyton'', is also placed outside the tracheophyte clade, as it did not possess true vascular tissue (in particular did not have tracheids), although its conducting tissue is more complex th ...
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Cladogram
A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to descendants, nor does it show how much they have changed, so many differing evolutionary trees can be consistent with the same cladogram. A cladogram uses lines that branch off in different directions ending at a clade, a group of organisms with a last common ancestor. There are many shapes of cladograms but they all have lines that branch off from other lines. The lines can be traced back to where they branch off. These branching off points represent a hypothetical ancestor (not an actual entity) which can be inferred to exhibit the traits shared among the terminal taxa above it. This hypothetical ancestor might then provide clues about the order of evolution of various features, adaptation, and other evolutionary narratives about ance ...
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