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Spinosaurinae
The Spinosauridae (or spinosaurids) are a clade or family of tetanuran theropod dinosaurs comprising ten to seventeen known genera. They came into prominence during the Cretaceous period. Spinosaurid fossils have been recovered worldwide, including Africa, Europe, South America and Asia. Their remains have generally been attributed to the Early to Mid Cretaceous. Spinosaurids were large bipedal carnivores. Their crocodilian-like skulls were long, low and narrow, bearing conical teeth with reduced or absent serrations. The tips of their upper and lower jaws fanned out into a spoon-shaped structure similar to a rosette, behind which there was a notch in the upper jaw that the expanded tip of the lower jaw fit into. The nostrils of spinosaurids were retracted to a position further back on the head than in most other theropods, and they had bony crests on their heads along the midline of their skulls. Their robust shoulders wielded stocky forelimbs, with three-fingered hands that ...
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Spinosaurus Aegyptiacus
''Spinosaurus'' (; ) is a genus of spinosaurid dinosaur that lived in what now is North Africa during the Cenomanian to upper Turonian stages of the Late Cretaceous period, about 99 to 93.5 million years ago. The genus was known first from Egyptian remains discovered in 1912 and described by German palaeontologist Ernst Stromer in 1915. The original remains were destroyed in World War II, but additional material came to light in the early 21st century. It is unclear whether one or two species are represented in the fossils reported in the scientific literature. The best known species is ''S. aegyptiacus'' from Egypt, although a potential second species, ''S. maroccanus'', has been recovered from Morocco. The contemporary spinosaurid genus ''Sigilmassasaurus'' has also been synonymized by some authors with ''S. aegyptiacus'', though other researchers propose it to be a distinct taxon. Another possible junior synonym is ''Oxalaia'' from the Alcântara Formation in Brazil. ' ...
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Spinosaurus
''Spinosaurus'' (; ) is a genus of spinosaurid dinosaur that lived in what now is North Africa during the Cenomanian to upper Turonian stages of the Late Cretaceous period, about 99 to 93.5 million years ago. The genus was known first from Egyptian remains discovered in 1912 and described by German palaeontologist Ernst Stromer in 1915. The original remains were destroyed in World War II, but additional material came to light in the early 21st century. It is unclear whether one or two species are represented in the fossils reported in the scientific literature. The best known species is ''S. aegyptiacus'' from Egypt, although a potential second species, ''S. maroccanus'', has been recovered from Morocco. The contemporary spinosaurid genus ''Sigilmassasaurus'' has also been synonymized by some authors with ''S. aegyptiacus'', though other researchers propose it to be a distinct taxon. Another possible junior synonym is ''Oxalaia'' from the Alcântara Formation in Brazil. ' ...
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Siamosaurus
''Siamosaurus'' (meaning "Siam reptile") is a genus of spinosaurid dinosaur that lived in what is now known as China and Thailand during the Early Cretaceous period (Barremian to Aptian) and is the first reported spinosaurid from Asia. It is confidently known only from tooth fossils; the first were found in the Sao Khua Formation, with more teeth later recovered from the younger Khok Kruat Formation. The only species ''Siamosaurus suteethorni'', whose name honours Thai palaeontologist Varavudh Suteethorn, was formally described in 1986. In 2009, four teeth from China previously attributed to a pliosaur—under the species "''Sinopliosaurus''" ''fusuiensis''—were identified as those of a spinosaurid, possibly ''Siamosaurus''. It is yet to be determined if two partial spinosaurid skeletons from Thailand and an isolated tooth from Japan also belong to ''Siamosaurus''. Since it is based only on teeth, ''Siamosaurus'' body size is uncertain, though it has been estimated at betw ...
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Baryonychinae
Baryonychinae is an extinct clade or subfamily of spinosaurids from the Early Cretaceous (Valanginian-Albian) of Britain, Portugal, and Niger. In 2021, it consisted of six genera: ''Ceratosuchops'', ''Cristatusaurus'', ''Riparovenator'', ''Suchomimus'', '' Suchosaurus'', and ''Baryonyx'', the nominal genus. The clade was named by Charig & Milner in 1986 and defined by Sereno ''et al''. in 1998 and Holtz ''et al.'' in 2004 as all taxa more closely related to ''Baryonyx walkeri'' than to ''Spinosaurus aegyptiacus''. Baryonychines were large, bipedal predators with elongated, crocodile-like skulls and lower jaw tips fanning out into rosettes bearing conical, often unserrated, teeth, and a distinct premaxillary notch. They possessed robust forelimbs supporting three-fingered hands with an enlarged first digit claw, to which the subfamily name indirectly refers. Members of this group, unlike the more derived Spinosaurinae, sported only low sails or none at all. History of discovery ...
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Vallibonavenatrix
''Vallibonavenatrix'' (meaning "Vallibona huntress" after the town near where its remains were found) is a genus of spinosaurid dinosaur from the Early Cretaceous (Barremian) Arcillas de Morella Formation of Castellón, Spain. The type and only species is ''Vallibonavenatrix cani'', known from a partial skeleton. Discovery and naming During the late 1980s and early 1990s, amateur fossil collector Juan Cano Forner was recovering bones from various localities in Els Ports Natural Park, located in the Province of Castellón, Spain. In one of these—the Santa Águeda locality in the town of Vallibona—he excavated numerous vertebrate remains dating to the Mesozoic era, among which were dinosaur fossils. Forner housed these fossils in a private collection at Sant Mateu, which the Generalitat Valenciana acknowledged as a museographic collection in 1994. In 2007, the Spanish palaeontologist Fernando Gómez-Fernández and colleagues published a provisional description on the pel ...
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Baryonyx
''Baryonyx'' () is a genus of theropod dinosaur which lived in the Barremian stage of the Early Cretaceous period, about 130–125 million years ago. The first skeleton was discovered in 1983 in the Smokejack Clay Pit, of Surrey, England, in sediments of the Weald Clay Formation, and became the holotype specimen of ''Baryonyx walkeri'', named by palaeontologists Alan J. Charig and Angela C. Milner in 1986. The generic name, ''Baryonyx'', means "heavy claw" and alludes to the animal's very large claw on the first finger; the specific name, ''walkeri'', refers to its discoverer, amateur fossil collector William J. Walker. The holotype specimen is one of the most complete theropod skeletons from the UK (and remains the most complete Spinosaurid), and its discovery attracted media attention. Specimens later discovered in other parts of the United Kingdom and Iberia have also been assigned to the genus, though many have since been moved to new genera. The holotype specimen, whic ...
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Irritator
''Irritator'' is a genus of spinosaurid dinosaur that lived in what is now Brazil during the Albian stage of the Early Cretaceous Period, about 113 to 110 million years ago. It is known from a nearly complete skull found in the Romualdo Formation of the Araripe Basin. Fossil dealers had acquired this skull and sold it to the State Museum of Natural History Stuttgart. In 1996, the specimen became the holotype of the type species ''Irritator challengeri''. The genus name comes from the word "irritation", reflecting the feelings of paleontologists who found the skull had been heavily damaged and altered by the collectors. The species name is a homage to the fictional character Professor Challenger from Arthur Conan Doyle's novels. Some paleontologists regard ''Angaturama limai''—known from a snout tip that was described later in 1996—as a potential junior synonym of ''Irritator''. Both animals hail from the same stratigraphic units of the Araripe Basin. It was also previousl ...
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Ichthyovenator
''Ichthyovenator'' is a genus of spinosaurid dinosaur that lived in what is now Laos, sometime between 125 and 113 million years ago, during the Aptian stage of the Early Cretaceous period. It is known from fossils collected from the Grès supérieurs Formation of the Savannakhet Basin, the first of which were found in 2010, consisting of a partial skeleton without the skull or limbs. This specimen became the holotype of the new genus and species ''Ichthyovenator laosensis'', and was described by palaeontologist Ronan Allain and colleagues in 2012. The generic name, meaning "fish hunter", refers to its assumed piscivorous lifestyle, while the specific name alludes to the country of Laos. In 2014, it was announced that more remains from the dig site had been recovered; these fossils included teeth, more vertebrae (backbones) and a pubic bone from the same individual. The holotype specimen is estimated to have been between long and to have weighed . ''Ichthyovenator''s te ...
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Iberospinus
''Iberospinus'' or (meaning " Iberian spine") is an extinct genus of spinosaurid dinosaur from the Early Cretaceous (Barremian) Papo Seco Formation of Portugal. The genus contains a single species, ''I. natarioi'', known from several assorted bones belonging to one individual. ''Iberospinus'' represents one of three known spinosaurid taxa from the Iberian Peninsula, the others being ''Vallibonavenatrix'', and ''Camarillasaurus''. It is important for its implications of the geographical origin of Spinosauridae and the suggested presence of an at least semi-aquatic lifestyle early in the evolution of this clade. Discovery and naming The first fossil material was discovered in 1999, with additional expeditions from 2004 to 2008. After being described as a specimen of ''Baryonyx'' in 2011, it was realised to have been a unique species in 2019. Additional material was discovered in a June 2020 expedition, after which ''Iberospinus'' was described as a new genus and species in 2022 ...
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Cristatusaurus
''Cristatusaurus'' is a genus of theropod dinosaur that lived during the Early Cretaceous Period of what is now Niger, 112 million years ago. It was a baryonychine member of the Spinosauridae, a group of large bipedal carnivores with well-built forelimbs and elongated, crocodile-like skulls. The type species ''Cristatusaurus lapparenti'' was named in 1998 by scientists Philippe Taquet and Dale Russell, on the basis of jaw bones and some vertebrae. Two claw fossils were also later assigned to ''Cristatusaurus''. The animal's generic name, which means "crested reptile", alludes to a sagittal crest on top of its snout; while the specific name is in honor of the French paleontologist Albert-Félix de Lapparent. ''Cristatusaurus'' is known from the Albian to Aptian Elrhaz Formation, where it would have coexisted with sauropod and iguanodontian dinosaurs, other theropods, and various crocodylomorphs. Originally proposed to be an indeterminate species of ''Baryonyx'', the identity of ...
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Oxalaia
''Oxalaia'' (in reference to the African deity '' Oxalá'') is a genus of spinosaurid dinosaur that lived in what is now the Northeast Region of Brazil during the Cenomanian stage of the Late Cretaceous period, sometime between 100.5 and 93.9 million years ago. Its only known fossils were found in 1999 on Cajual Island in the rocks of the Alcântara Formation, which is known for its abundance of fragmentary, isolated fossil specimens. The remains of ''Oxalaia'' were described in 2011 by Brazilian palaeontologist Alexander Kellner and colleagues, who assigned the specimens to a new genus containing one species, ''Oxalaia quilombensis''. The species name refers to the Brazilian quilombo settlements. ''Oxalaia quilombensis'' is the eighth officially named theropod species from Brazil and the largest carnivorous dinosaur discovered there. It is closely related to the African genus ''Spinosaurus'', and/or may be a junior synonym of this taxon. Although ''Oxalaia'' is known only from ...
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Suchomimus
''Suchomimus'' (meaning "crocodile mimic") is a genus of spinosaurid dinosaur that lived between 125 and 112 million years ago in what is now Niger, during the Aptian to early Albian stages of the Early Cretaceous period. It was named and described by palaeontologist Paul Sereno and colleagues in 1998, based on a partial skeleton from the Elrhaz Formation. ''Suchomimus'''s long and shallow skull, similar to that of a crocodile, earns it its generic name, while the specific name ''Suchomimus tenerensis'' alludes to the locality of its first remains, the Ténéré Desert. ''Suchomimus'' was a relatively large theropod, reaching in length and weighing . However, the age of the holotype specimen is uncertain, so it is unclear whether this size estimate would have been its maximum. The narrow skull of ''Suchomimus'' was perched on a short neck, and its forelimbs were powerfully built, bearing a giant claw on each thumb. Along the midline of the animal's back ran a low dorsal ...
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