Iberospinus
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Iberospinus
''Iberospinus'' or (meaning " Iberian spine") is an extinct genus of spinosaurid dinosaur from the Early Cretaceous (Barremian) Papo Seco Formation of Portugal. The genus contains a single species, ''I. natarioi'', known from several assorted bones belonging to one individual. ''Iberospinus'' represents one of three known spinosaurid taxa from the Iberian Peninsula, the others being ''Vallibonavenatrix'', and ''Camarillasaurus''. It is important for its implications of the geographical origin of Spinosauridae and the suggested presence of an at least semi-aquatic lifestyle early in the evolution of this clade. Discovery and naming The first fossil material was discovered in 1999, with additional expeditions from 2004 to 2008. After being described as a specimen of ''Baryonyx'' in 2011, it was realised to have been a unique species in 2019. Additional material was discovered in a June 2020 expedition, after which ''Iberospinus'' was described as a new genus and species in 2022 ...
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Iberospinus Scapula
''Iberospinus'' or (meaning " Iberian spine") is an extinct genus of spinosaurid dinosaur from the Early Cretaceous (Barremian) Papo Seco Formation of Portugal. The genus contains a single species, ''I. natarioi'', known from several assorted bones belonging to one individual. ''Iberospinus'' represents one of three known spinosaurid taxa from the Iberian Peninsula, the others being ''Vallibonavenatrix'', and ''Camarillasaurus''. It is important for its implications of the geographical origin of Spinosauridae and the suggested presence of an at least semi-aquatic lifestyle early in the evolution of this clade. Discovery and naming The first fossil material was discovered in 1999, with additional expeditions from 2004 to 2008. After being described as a specimen of ''Baryonyx'' in 2011, it was realised to have been a unique species in 2019. Additional material was discovered in a June 2020 expedition, after which ''Iberospinus'' was described as a new genus and species in 2022 ...
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Iberospinus CT-scan
''Iberospinus'' or (meaning " Iberian spine") is an extinct genus of spinosaurid dinosaur from the Early Cretaceous (Barremian) Papo Seco Formation of Portugal. The genus contains a single species, ''I. natarioi'', known from several assorted bones belonging to one individual. ''Iberospinus'' represents one of three known spinosaurid taxa from the Iberian Peninsula, the others being ''Vallibonavenatrix'', and ''Camarillasaurus''. It is important for its implications of the geographical origin of Spinosauridae and the suggested presence of an at least semi-aquatic lifestyle early in the evolution of this clade. Discovery and naming The first fossil material was discovered in 1999, with additional expeditions from 2004 to 2008. After being described as a specimen of ''Baryonyx'' in 2011, it was realised to have been a unique species in 2019. Additional material was discovered in a June 2020 expedition, after which ''Iberospinus'' was described as a new genus and species in 2022 ...
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Iberospinus Life Reconstruction
''Iberospinus'' or (meaning " Iberian spine") is an extinct genus of spinosaurid dinosaur from the Early Cretaceous (Barremian) Papo Seco Formation of Portugal. The genus contains a single species, ''I. natarioi'', known from several assorted bones belonging to one individual. ''Iberospinus'' represents one of three known spinosaurid taxa from the Iberian Peninsula, the others being ''Vallibonavenatrix'', and ''Camarillasaurus''. It is important for its implications of the geographical origin of Spinosauridae and the suggested presence of an at least semi-aquatic lifestyle early in the evolution of this clade. Discovery and naming The first fossil material was discovered in 1999, with additional expeditions from 2004 to 2008. After being described as a specimen of ''Baryonyx'' in 2011, it was realised to have been a unique species in 2019. Additional material was discovered in a June 2020 expedition, after which ''Iberospinus'' was described as a new genus and species in 2022 ...
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Spinosauridae
The Spinosauridae (or spinosaurids) are a clade or family of tetanuran theropod dinosaurs comprising ten to seventeen known genera. They came into prominence during the Cretaceous period. Spinosaurid fossils have been recovered worldwide, including Africa, Europe, South America and Asia. Their remains have generally been attributed to the Early to Mid Cretaceous. Spinosaurids were large bipedal carnivores. Their crocodilian-like skulls were long, low and narrow, bearing conical teeth with reduced or absent serrations. The tips of their upper and lower jaws fanned out into a spoon-shaped structure similar to a rosette, behind which there was a notch in the upper jaw that the expanded tip of the lower jaw fit into. The nostrils of spinosaurids were retracted to a position further back on the head than in most other theropods, and they had bony crests on their heads along the midline of their skulls. Their robust shoulders wielded stocky forelimbs, with three-fingered hands that b ...
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Octávio Mateus
Octávio Mateus (born 1975) is a Portuguese dinosaur paleontologist and biologist Professor of Paleontology at the Faculdade de Ciências e Tecnologia da Universidade Nova de Lisboa. He graduated in Universidade de Évora and received his PhD at Universidade Nova de Lisboa in 2005. He collaborates with Museu da Lourinhã, known for their dinosaur collection. His former PhD advisor was Miguel Telles Antunes. He is an expert on dinosaurs, particularly Late Jurassic dinosaurs of Portugal, but he has also worked with specimens from Angola, the USA, Greenland, China and Morocco. New dinosaur taxa he has helped name include '' Lourinhanosaurus antunesi'' (1998), '' Dinheirosaurus lourinhanensis'' (1999), '' Tangvayosaurus hoffeti'' (1999), '' Draconyx loureiroi'' (2001), '' Lusotitan atalaiensis'' (2003), '' Europasaurus holgeri'' (2006), '' Allosaurus europaeus'' (2006), '' Angolatitan adamastor'' (2011)'','' ''Torvosaurus gurneyi'' (2014), ''Zby atlanticus'' (2014), ''Galeamopus'' (20 ...
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Baryonyx
''Baryonyx'' () is a genus of theropod dinosaur which lived in the Barremian stage of the Early Cretaceous period, about 130–125 million years ago. The first skeleton was discovered in 1983 in the Smokejack Clay Pit, of Surrey, England, in sediments of the Weald Clay Formation, and became the holotype specimen of ''Baryonyx walkeri'', named by palaeontologists Alan J. Charig and Angela C. Milner in 1986. The generic name, ''Baryonyx'', means "heavy claw" and alludes to the animal's very large claw on the first finger; the specific name, ''walkeri'', refers to its discoverer, amateur fossil collector William J. Walker. The holotype specimen is one of the most complete theropod skeletons from the UK (and remains the most complete Spinosaurid), and its discovery attracted media attention. Specimens later discovered in other parts of the United Kingdom and Iberia have also been assigned to the genus, though many have since been moved to new genera. The holotype specimen, whic ...
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2022 In Archosaur Paleontology
This article records new taxa of fossil archosaurs of every kind that are scheduled described during the year 2022, as well as other significant discoveries and events related to paleontology of archosaurs that are scheduled to occur in the year 2022. Pseudosuchians New pseudosuchian taxa General pseudosuchian research * A study on the mandible embryogenesis in extant caimans, and on its implications for the knowledge of the evolution of postdentary lower jaw of pseudosuchians, is published by Bona ''et al.'' (2022). * A study on the musculature of crocodylian and fossil suchian jaws, investigating the impact of the flattening of the skulls of suchians in their evolutionary history on their muscle anatomy, is published by Sellers ''et al.'' (2022). * Revision of ''Tsylmosuchus donensis'' and ''Scythosuchus basileus'' is published by Sennikov (2022), who interprets the latter taxon as a junior synonym of the former one, and interprets ''T. donensis'' as a likely member of ...
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Papo Seco Formation
The Papo Seco Formation is a geological formation in Portugal, whose strata date back to the Early Cretaceous. Dinosaur fossils are among the fossils that have been recovered from the formation.Weishampel et al., 2004, p.562Papo Seco Formation
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Name

The unit was previously referred to as the "Gres a Dinosauriens", due to the abundant dinosaur remains found in the past.Aillud, 2001, p.211


Description

The Papo Seco Formation (Rey, 1992) marks the return to

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Vallibonavenatrix
''Vallibonavenatrix'' (meaning "Vallibona huntress" after the town near where its remains were found) is a genus of spinosaurid dinosaur from the Early Cretaceous (Barremian) Arcillas de Morella Formation of Castellón, Spain. The type and only species is ''Vallibonavenatrix cani'', known from a partial skeleton. Discovery and naming During the late 1980s and early 1990s, amateur fossil collector Juan Cano Forner was recovering bones from various localities in Els Ports Natural Park, located in the Province of Castellón, Spain. In one of these—the Santa Águeda locality in the town of Vallibona—he excavated numerous vertebrate remains dating to the Mesozoic era, among which were dinosaur fossils. Forner housed these fossils in a private collection at Sant Mateu, which the Generalitat Valenciana acknowledged as a museographic collection in 1994. In 2007, the Spanish palaeontologist Fernando Gómez-Fernández and colleagues published a provisional description on the pel ...
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Camarillasaurus
''Camarillasaurus'' (meaning " Camarillas lizard") is a genus of spinosaurid dinosaur from the Early Cretaceous period (Barremian) of Camarillas, Teruel Province, in what is now northeastern Spain. Described in 2014, it was originally identified as a ceratosaurian theropod, before being referred to the Spinosauridae in 2021. History of research Fossils of ''Camarillasaurus'' were discovered in the Camarillas Formation. The type species, ''Camarillasaurus cirugedae'', was described by palaeontologists Bárbara Sánchez-Hernández and Michael J. Benton. The describers considered it to be a basal ceratosaur, somewhat unusual given its young age. However, in an abstract given at the EAVP 2019 conference, Oliver Rauhut and colleagues suggested it is a member of the Spinosauridae rather than a ceratosaur, based on characters of the posterior caudal vertebrae and newly excavated material at the type locality; a chapter of a 2019 dissertation by Adun Samathi concurred with that conclu ...
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Calcaneus
In humans and many other primates, the calcaneus (; from the Latin ''calcaneus'' or ''calcaneum'', meaning heel) or heel bone is a bone of the tarsus of the foot which constitutes the heel. In some other animals, it is the point of the hock. Structure In humans, the calcaneus is the largest of the tarsal bones and the largest bone of the foot. Its long axis is pointed forwards and laterally. The talus bone, calcaneus, and navicular bone are considered the proximal row of tarsal bones. In the calcaneus, several important structures can be distinguished:Platzer (2004), p 216 There is a large calcaneal tuberosity located posteriorly on plantar surface with medial and lateral tubercles on its surface. Besides, there is another peroneal tubecle on its lateral surface. On its lower edge on either side are its lateral and medial processes (serving as the origins of the abductor hallucis and abductor digiti minimi). The Achilles tendon is inserted into a roughened area on its superio ...
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Ungual
An ungual (from Latin ''unguis'', i.e. ''nail'') is a highly modified distal toe bone which ends in a hoof, claw, or nail. Elephants and ungulates have ungual phalanges, as did the sauropods and horned dinosaurs. A claw is a highly modified ungual phalanx. As an adjective, ungual means ''related to nail Nail or Nails may refer to: In biology * Nail (anatomy), toughened protective protein-keratin (known as alpha-keratin, also found in hair) at the end of an animal digit, such as fingernail * Nail (beak), a plate of hard horny tissue at the tip ...'', as in ''periungual'' (around the nail). References External links Mammal anatomy {{animal-anatomy-stub ...
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