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Solemydidae
The Helochelydridae are an extinct family of stem-turtles known from fossils found in North America and Europe spanning the Early to Late Cretaceous. Description The skull, shell and osteoderms of helochelydrids are covered in small, cylindrical protuberances, which are a distinctive characteristic of the group. They are thought to be terrestrial, based on the presence of limb osteoderms and bone histology. Their skull morphology is dissimilar to that of extant tortoises, suggesting an omnivorous habit similar to that of box turtles. Taxonomy Helochelydridae includes all turtles that closely related to '' Helochelydra'' than '' Sichuanchelys'', '' Meiolania'', or extant turtles. Although referred to as Solemydidae in recent literature on extinct turtles, Helochelydridae has priority over Solemydidae. They are placed as part of the clade Perichelydia. Some recent studies have recovered them as paracryptodires, though other studies have found them to be more basal than pa ...
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Helochelydra
''Helochelydra'' is an extinct genus of extinct stem turtle known from the Early Cretaceous (Barremian) of the Isle of Wight, southern England. Phylogeny ''Helochelydra'' is a member of the stem turtle family Helochelydridae, which is known from Late Jurassic to Late Cretaceous deposits in North America and Europe. Cladistic analysis recovers Helochelydridae outside the clade leading to crown turtles (Testudines).W. G. Joyce, S. D. Chapman, R. T. J. Moody, C. A. Walker (2011) The skull of the solemydid turtle ''Helochelydra nopcsai'' from the Early Cretaceous of the Isle of Wight (UK) and a review of Solemydidae. ''Special Papers in Palaeontology'': 75-97. Taxonomy ''Helochelydra'' was named in 1928 by Franz Baron Nopcsa for a partial shell ( NHMUK R171) from the Early Cretaceous Wessex Formation of the Isle of Wight that Lydekker (1889) had referred to ''Tretosternon punctatum'', a turtle taxon from the Purbeck Group of Dorset, but no species name was provided. Lapparent de Bro ...
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Solemys
''Solemys'' is an extinct genus of stem turtle known from the Late Cretaceous The Late Cretaceous (100.5–66 Ma) is the younger of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''creta'', the ... (late Campanian-early Maastrichtian) of southern France and eastern Spain. Taxonomy Two species are known, the type species ''Solemys gaudryi'' (Matheron, 1869) and ''S. vermiculata'' de Lapparent de Broin and Murelaga, 1996. The former was originally assigned to the Cenozoic turtle genus ''Apholidemys'' by Matheron (1869), but de Lapparent de Broin and Murelaga (1996) recognized it as a stem turtle distinct from ''Apholidemys'' and renamed it ''Solemys''.F. d.e. Lapparent de Broin and X. Murelaga. 1996. Une nouvelle faune de chéloniens dans le Crétacé supérieur européen. Comptes Rendus de l’Académie des Sciences de Paris 323:729-735 References ...
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Helochelys
''Helochelys'' is an extinct Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and ... genus of stem turtle known from the Late Cretaceous (Cenomanian) of southern Germany. References Fossil taxa described in 1854 Early Cretaceous turtles Extinct turtles Testudinata {{Paleo-turtle-stub ...
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Purbeck Group
The Purbeck Group is an Upper Jurassic to Lower Cretaceous lithostratigraphic group (a sequence of rock strata) in south-east England. The name is derived from the district known as the Isle of Purbeck in Dorset where the strata are exposed in the cliffs west of Swanage. The Purbeck Group is famous for its fossils of reptiles and early mammals. This sequence of rocks has gone by various names in the past including amongst others the Purbeck Beds, Purbeck Formation, Purbeck Limestone Formation and Purbeck Stone. Rocks of this age have in the past been called the Purbeckian stage by European geologists. The Purbeckian corresponds with the Tithonian to Berriasian stages of the internationally used geologic timescale. Outcrops The Purbeck Group outcrops follow the line of the Jurassic outcrop from Dorset, through the Vale of Wardour, Swindon, Garsington, Brill and Aylesbury. In East Sussex, the Purbeck Group outcrops at three locations north and northwest west of Battle, East Suss ...
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Angeac-Charente Bonebed
The Angeac-Charente bonebed is a fossil deposit located near Angeac-Charente in western France. It dates to the Berriasian stage of the Early Cretaceous, and is coeval with the Purbeck Group of Southern England. It has amongst the most diverse assemblages of earliest Cretaceous vertebrates known from Europe. History of discovery Dinosaur bones were first found at the site in 2008. The fossils were exposed during quarrying for overlying Pleistocene aged sand and gravel. After more bones were discovered in 2010, a team was set up composed of people from Musée d’Angoulême, Rennes University and the Muséum national d’Histoire naturelle to excavate the site. Due the promising finds, since 2011 excavations have been conducted at the site annually.Ronan Allain, Romain Vullo, Lee Rozada, Jérémy Anquetin, Renaud Bourgeais, et al.Vertebrate paleobiodiversity of the Early Cretaceous (Berriasian) Angeac-Charente Lagerstätte (southwestern France): implications for continental faunal ...
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Maastrichtian
The Maastrichtian () is, in the ICS geologic timescale, the latest age (uppermost stage) of the Late Cretaceous Epoch or Upper Cretaceous Series, the Cretaceous Period or System, and of the Mesozoic Era or Erathem. It spanned the interval from . The Maastrichtian was preceded by the Campanian and succeeded by the Danian (part of the Paleogene and Paleocene). The Cretaceous–Paleogene extinction event (formerly known as the Cretaceous–Tertiary extinction event) occurred at the end of this age. In this mass extinction, many commonly recognized groups such as non-avian dinosaurs, plesiosaurs and mosasaurs, as well as many other lesser-known groups, died out. The cause of the extinction is most commonly linked to an asteroid about wide colliding with Earth, ending the Cretaceous. Stratigraphic definitions Definition The Maastrichtian was introduced into scientific literature by Belgian geologist André Hubert Dumont in 1849, after studying rock strata of the Chalk Group c ...
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Campanian
The Campanian is the fifth of six ages of the Late Cretaceous Epoch on the geologic timescale of the International Commission on Stratigraphy (ICS). In chronostratigraphy, it is the fifth of six stages in the Upper Cretaceous Series. Campanian spans the time from 83.6 (± 0.2) to 72.1 (± 0.2) million years ago. It is preceded by the Santonian and it is followed by the Maastrichtian. The Campanian was an age when a worldwide sea level rise covered many coastal areas. The morphology of some of these areas has been preserved: it is an unconformity beneath a cover of marine sedimentary rocks. Etymology The Campanian was introduced in scientific literature by Henri Coquand in 1857. It is named after the French village of Champagne in the department of Charente-Maritime. The original type locality was a series of outcrop near the village of Aubeterre-sur-Dronne in the same region. Definition The base of the Campanian Stage is defined as a place in the stratigraphic column wher ...
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Upper Greensand Formation
The Upper Greensand Formation is a Cretaceous formation of Albian to Cenomanian in age, found within the Wessex Basin and parts of the Weald Basin in southern England. It overlies the Gault Clay and underlies the Chalk Group. It varies in thickness from zero to 75 m. It is predominantly a glauconitic fine-grained sandstone, locally becoming silty. Fragmentary dinosaur remains, such as those assigned to ''Iuticosaurus'', have been recovered from this formation. It has been quarried as a building stone from Roman times, and used in London and the area of its outcrop from Devon to East Sussex. Use Sandstones from the Upper Greensand have been used as building stone since at least Roman times. Reigate Stone was mined from the Upper Greensand in north east Surrey throughout the middle ages and early modern period, forming the most important source of freestone in Medieval London. In Dorset, the Shaftesbury Sandstone was quarried between Shaftesbury and Okeford Fitzpaine and used i ...
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Cambridge Greensand
The Cambridge Greensand is a geological unit in England whose strata are earliest Cenomanian in age. It lies above the erosive contact between the Gault Formation and the Chalk Group in the vicinity of Cambridgeshire, and technically forms the lowest member bed of the West Melbury Marly Chalk Formation.Cambridge Greensand
at BGS
It is a remanié deposit, containing reworked fossils of late age, including those of dinosaurs and pterosaurs.


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The lithology is made out of

Cenomanian
The Cenomanian is, in the ICS' geological timescale, the oldest or earliest age of the Late Cretaceous Epoch or the lowest stage of the Upper Cretaceous Series. An age is a unit of geochronology; it is a unit of time; the stage is a unit in the stratigraphic column deposited during the corresponding age. Both age and stage bear the same name. As a unit of geologic time measure, the Cenomanian Age spans the time between 100.5 and 93.9 million years ago (Mya). In the geologic timescale, it is preceded by the Albian and is followed by the Turonian. The Upper Cenomanian starts around at 95 Mya. The Cenomanian is coeval with the Woodbinian of the regional timescale of the Gulf of Mexico and the early part of the Eaglefordian of the regional timescale of the East Coast of the United States. At the end of the Cenomanian, an anoxic event took place, called the Cenomanian-Turonian boundary event or the "Bonarelli event", that is associated with a minor extinction event for marine spec ...
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Grünsandstein Formation
The Grünsandstein Formation is a geologic formation in Germany. It preserves fossils dating back to the Cretaceous period. See also * List of fossiliferous stratigraphic units in Germany See also * Lists of fossiliferous stratigraphic units in Europe These lists of fossiliferous stratigraphic units in Europe enumerate the rock layers which preserve the fossilized remains of ancient life in Europe by the modern countries wher ... References * Cretaceous Germany {{Cretaceous-stub ...
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