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Sauriurae
Sauriurae (meaning "lizard tails" in Greek) is a now-deprecated subclass of birds created by Ernst Haeckel in 1866. It was intended to include '' Archaeopteryx'' and distinguish it from all other birds then known, which he grouped in the sister-group Ornithurae ("bird tails"). The distinction Haeckel referred to in this name is that ''Archaeopteryx'' possesses a long, reptile-like tail, while all other birds known to him had short tails with few vertebrae, fused at the end into a pygostyle.Haeckel, Ernst. (1866) "Generelle Morphologie der Organismen" Berlin: Georg Reimer. 462 pp. The unit was not much referred to, and when Hans Friedrich Gadow in 1893 erected Archaeornithes for basically the same fossils, this became the common name for the early reptile-like grade of birds. Ji Qiang and Larry Martin have continued to refer to the Sauriurae as a valid natural group.Ji, Q., and Ji, S. (1996). "On discovery of the earliest bird fossil in China and the origin of birds." ''Chinese ...
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Archaeornithes
The Archaeornithes, classically Archæornithes, is an extinct group of the first primitive, reptile-like birds. It is an evolutionary grade of transitional fossils, the primitive birds halfway between non avian dinosaur ancestors and the derived modern birds (avian dinosaur). Fossils of early birds were poorly known until the late 20th century. Of those known, all fell into either the relatively modernly built birds with fused ribcage and the breastbone extended into a keel, or the " Urvogels" of the Solnhofen Plattenkalk of late Jurassic age. As the physiological and anatomical difference between the two was so great, the subclass Archaeornithes was erected for the latter. With the unearthing of several well preserved early bird fossils in the last decades of the 20th century and early 21st century, our knowledge of the evolution of birds has increased dramatically. The evolution of the modern avian traits such as the compact body, clawless wing and the alula are now known to ...
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Ornithurae
Ornithurae (meaning "bird tails" in Greek) is a natural group which includes the common ancestor of ''Ichthyornis'', ''Hesperornis'', and all modern birds as well as all other descendants of that common ancestor. Classification Ernst Haeckel coined the name in 1866 and included in the group all "true birds" with the "characteristic tail morphology of all extant birds" (translation by Jacques Gauthier). This distinguishes the group from ''Archaeopteryx'', which Haeckel placed in another new group called Sauriurae. Said simply, modern birds have short, fused pygostyle tails, while ''Archaeopteryx'' retained the long tail characteristic of non-avian theropod dinosaurs.Haeckel, Ernst (1866). ''Generelle Morphologie der Organismen''. Berlin: Georg Reimer. Gauthier converted Ornithurae into a clade, giving it a branch-based definition: "extant birds and all other taxa, such as ''Ichthyornis'' and Hesperornithes, that are closer to extant birds than is ''Archaeopteryx''". Later he and ...
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Pygostyle
Pygostyle describes a skeletal condition in which the final few caudal vertebrae are fused into a single ossification, supporting the tail feathers and musculature. In modern birds, the rectrices attach to these. The pygostyle is the main component of the uropygium, a structure colloquially known as the bishop's nose, parson's nose, pope's nose, or sultan's nose. This is the fleshy protuberance visible at the posterior end of a bird (most commonly a chicken or turkey) that has been dressed for cooking. It has a swollen appearance because it also contains the uropygial gland that produces preen oil. Evolution Pygostyles probably began to evolve very early in the Cretaceous period, perhaps 140 – 130 million years ago. The earliest known species to have evolved a pygostyle were members of the Confuciusornithidae. The structure provided an evolutionary advantage, as a completely mobile tail as found in species such as '' Archaeopteryx'' is detrimental to its use for flight contro ...
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Evolutionary Grade
A grade is a taxon united by a level of morphological or physiological complexity. The term was coined by British biologist Julian Huxley, to contrast with clade, a strictly phylogenetic unit. Definition An evolutionary grade is a group of species united by morphological or physiological traits, that has given rise to another group that has major differences from the ancestral condition, and is thus not considered part of the ancestral group, while still having enough similarities that we can group them under the same clade. The ancestral group will not be phylogenetically complete (i.e. will not form a clade), so will represent a paraphyletic taxon. In order to fully understand evolutionary grades, one must first get a better understanding of Phylogenetics, defined as "''In biology, is the study of the evolutionary history and relationships among individuals or groups of organisms (e.g. species, or populations). These relationships are discovered through phylogenetic i ...
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Vertebrate Subclasses
Vertebrates () comprise all animal taxa within the subphylum Vertebrata () (chordates with backbones), including all mammals, birds, reptiles, amphibians, and fish. Vertebrates represent the overwhelming majority of the phylum Chordata, with currently about 69,963 species described. Vertebrates comprise such groups as the following: * jawless fish, which include hagfish and lampreys * jawed vertebrates, which include: ** cartilaginous fish (sharks, rays, and ratfish) ** bony vertebrates, which include: *** ray-fins (the majority of living bony fish) *** lobe-fins, which include: **** coelacanths and lungfish **** tetrapods (limbed vertebrates) Extant vertebrates range in size from the frog species ''Paedophryne amauensis'', at as little as , to the blue whale, at up to . Vertebrates make up less than five percent of all described animal species; the rest are invertebrates, which lack vertebral columns. The vertebrates traditionally include the hagfish, which do not have pro ...
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Phylogenetic Nomenclature
Phylogenetic nomenclature is a method of nomenclature for taxa in biology that uses phylogenetic definitions for taxon names as explained below. This contrasts with the traditional approach, in which taxon names are defined by a '' type'', which can be a specimen or a taxon of lower rank, and a description in words. Phylogenetic nomenclature is currently regulated by the '' International Code of Phylogenetic Nomenclature'' (''PhyloCode''). Definitions Phylogenetic nomenclature ties names to clades, groups consisting of an ancestor and all its descendants. These groups can equivalently be called monophyletic. There are slightly different ways of specifying the ancestor, which are discussed below. Once the ancestor is specified, the meaning of the name is fixed: the ancestor and all organisms which are its descendants are included in the named taxon. Listing all these organisms (i.e. providing a full circumscription) requires the full phylogenetic tree to be known. In practice, ther ...
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Paraphyletic
In taxonomy (general), taxonomy, a group is paraphyletic if it consists of the group's most recent common ancestor, last common ancestor and most of its descendants, excluding a few Monophyly, monophyletic subgroups. The group is said to be paraphyletic ''with respect to'' the excluded subgroups. In contrast, a monophyletic group (a clade) includes a common ancestor and ''all'' of its descendants. The terms are commonly used in phylogenetics (a subfield of biology) and in the tree model of historical linguistics. Paraphyletic groups are identified by a combination of Synapomorphy and apomorphy, synapomorphies and symplesiomorphy, symplesiomorphies. If many subgroups are missing from the named group, it is said to be polyparaphyletic. The term was coined by Willi Hennig to apply to well-known taxa like Reptilia (reptiles) which, as commonly named and traditionally defined, is paraphyletic with respect to mammals and birds. Reptilia contains the last common ancestor of reptiles a ...
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Avialae
Avialae ("bird wings") is a clade containing the only living dinosaurs, the birds. It is usually defined as all theropod dinosaurs more closely related to birds (Aves) than to deinonychosaurs, though alternative definitions are occasionally used (see below). ''Archaeopteryx lithographica'', from the late Jurassic Period Solnhofen Formation of Germany, is possibly the earliest known avialan which may have had the capability of powered flight, though it might have been a deinonychosaur instead. Several older (but non flight-capable) avialans are known from the late Jurassic Tiaojishan Formation of China, dated to about 160 million years ago. Definition Most researchers define Avialae as branch-based clade, though definitions vary. Many authors have used a definition similar to "all theropods closer to birds than to ''Deinonychus''."Weishampel, David B.; Dodson, Peter; Osmólska, Halszka (eds.) (2004). ''The Dinosauria'', Second Edition. University of California Press., 861 pp. ...
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Julia Clarke
Julia Allison Clarke is an American paleontologist and evolutionary biologist who studies the evolution of birds and the dinosaurs most closely related to living birds. She is the John A. Wilson Professor in Vertebrate Paleontology in the Jackson School of Geosciences and a Howard Hughes Medical Institute Professor at the University of Texas at Austin. Education Clarke graduated with a B.A. in Comparative Literature and Geobiology from Brown University in 1995. She went on to study at Yale University, earning a Ph.D. from the Department of Geology and Geophysics in 2002. Career In 2005, Clarke led a research team that reexamined a fossil discovered within the rocks of Vega Island in the Antarctic in 1992. The use of computer tomography (CT) scans allowed for a new and more detailed analysis of the partial skeleton and it was determined to be ''Vegavis iaai,'' an extinct Antarctic bird and early relative of ducks and geese and thought to be the only species of modern bird to hav ...
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Ancient Greek
Ancient Greek includes the forms of the Greek language used in ancient Greece and the ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Dark Ages (), the Archaic period (), and the Classical period (). Ancient Greek was the language of Homer and of fifth-century Athenian historians, playwrights, and philosophers. It has contributed many words to English vocabulary and has been a standard subject of study in educational institutions of the Western world since the Renaissance. This article primarily contains information about the Epic and Classical periods of the language. From the Hellenistic period (), Ancient Greek was followed by Koine Greek, which is regarded as a separate historical stage, although its earliest form closely resembles Attic Greek and its latest form approaches Medieval Greek. There were several regional dialects of Ancient Greek, of which Attic Greek developed into Koine. Dia ...
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Reptile
Reptiles, as most commonly defined are the animals in the class Reptilia ( ), a paraphyletic grouping comprising all sauropsids except birds. Living reptiles comprise turtles, crocodilians, squamates (lizards and snakes) and rhynchocephalians (tuatara). As of March 2022, the Reptile Database includes about 11,700 species. In the traditional Linnaean classification system, birds are considered a separate class to reptiles. However, crocodilians are more closely related to birds than they are to other living reptiles, and so modern cladistic classification systems include birds within Reptilia, redefining the term as a clade. Other cladistic definitions abandon the term reptile altogether in favor of the clade Sauropsida, which refers to all amniotes more closely related to modern reptiles than to mammals. The study of the traditional reptile orders, historically combined with that of modern amphibians, is called herpetology. The earliest known proto-reptiles originated around ...
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Class (biology)
In biological classification, class ( la, classis) is a taxonomic rank, as well as a taxonomic unit, a taxon, in that rank. It is a group of related taxonomic orders. Other well-known ranks in descending order of size are life, domain, kingdom, phylum, order, family, genus, and species, with class fitting between phylum and order. History The class as a distinct rank of biological classification having its own distinctive name (and not just called a ''top-level genus'' ''(genus summum)'') was first introduced by the French botanist Joseph Pitton de Tournefort in his classification of plants that appeared in his ''Eléments de botanique'', 1694. Insofar as a general definition of a class is available, it has historically been conceived as embracing taxa that combine a distinct ''grade'' of organization—i.e. a 'level of complexity', measured in terms of how differentiated their organ systems are into distinct regions or sub-organs—with a distinct ''type'' of construction, ...
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