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Rhamphocephalus
''Rhamphocephalus'' ("beak head") is an extinct genus of fossil reptile from the Middle Jurassic (Bathonian stage) Great Oolite Group of Gloucestershire, England. The name was erected as a genus of pterosaur and became a 'wastebasket taxon' for British Jurassic pterosaur remains until a recent revision. ''Rhamphocephalus'' comprises several named species, two of which are pterosaurian, but the type species - ''R. prestwichi'' - is based on remains now identified as a thalattosuchian. Because it is poorly preserved and lacks features that distinguish it from other thalattosuchians, ''R. prestwichi'' is considered an invalid species and the genus ''Rhamphocephalus'' is a nomen dubium. Reassessments of other ''Rhamphocephalus'' species suggest they are also undiagnostic to species level, although they have properties allowing referral to some Jurassic pterosaur groups. Taxonomy The type species, ''R. prestwichi'', is known from the skull roof fragment OUM J.28266, found the Kineton T ...
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Dolicorhamphus
''Dolicorhamphus'' is an extinct genus of pterosaurs from the Middle Jurassic Taynton Limestone Formation and Fuller's Earth Formations of England. The genus contains two species, ''D. bucklandii'' and ''D. depressirostris''. Both species of ''Dolicorhamphus'' were assigned to the wastebasket genus '' Rhamphocephalus'', considered a "rhamphorhynchoid" pterosaur. However, the type species of ''Rhamphocephalus'', ''R. prestwichii'', was reinterpreted as a thalattosuchian by O'Sullivan and Martill on 2018, but ''"R." depressirostris'' and ''"R." bucklandii'' were clearly pterosaurian; the former was considered to be in the Scaphognathinae while the latter was only referred to a more general Rhamphorhynchidae. Andres (2021) resurrected the genus ''Dolicorhamphus'' for the two, finding it to be a valid genus of pterosaurs closely related to ''Klobiodon'', although he did not assign a type species. This generic assignment has since been followed by other researchers. ''D. bucklandi'' i ...
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Klobiodon
''Klobiodon'' is a genus of rhamphorhynchid pterosaur from the Middle Jurassic Taynton Limestone Formation of Oxfordshire, England. Etymology The type species of ''Klobiodon'' is ''Klobiodon rochei''. The generic name ''Klobiodon'' means "small cage tooth," from the Greek κλωβίον, ''klobion'', "little cage", and ὀδών, ''odon'', "tooth", in reference to the large anterior laniaries that appear to form a fish grab, while the specific name ''rochei'' honors the comic book artist Nick Roche for his anatomically correct designs inspired by dinosaurs. History The holotype of ''Klobiodon rochei'', NHMUK PV OR 47991, was first mentioned by George Robert Waterhouse (1878) as part of the collection of the British Museum of Natural History, where he mentions Richard Owen intended to name the specimen ''Pterodactylus raptor'' in an unpublished manuscript. It had been donated by Robert Marsham. Richard Lydekker (1888) referred the specimen to ''Rhamphorhynchus depressirostris'' ( ...
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Genus
Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus comes above species and below family (taxonomy), family. In binomial nomenclature, the genus name forms the first part of the binomial species name for each species within the genus. :E.g. ''Panthera leo'' (lion) and ''Panthera onca'' (jaguar) are two species within the genus ''Panthera''. ''Panthera'' is a genus within the family Felidae. The composition of a genus is determined by taxonomy (biology), taxonomists. The standards for genus classification are not strictly codified, so different authorities often produce different classifications for genera. There are some general practices used, however, including the idea that a newly defined genus should fulfill these three criteria to be descriptively useful: # monophyly – all descendants ...
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Middle Jurassic
The Middle Jurassic is the second epoch of the Jurassic Period. It lasted from about 174.1 to 163.5 million years ago. Fossils of land-dwelling animals, such as dinosaurs, from the Middle Jurassic are relatively rare, but geological formations containing land animal fossils include the Forest Marble Formation in England, the Kilmaluag Formation in Scotland,British Geological Survey. 2011Stratigraphic framework for the Middle Jurassic strata of Great Britain and the adjoining continental shelf: research report RR/11/06 British Geological Survey, Keyworth, Nottingham. the Daohugou Beds in China, the Itat Formation in Russia, and the Isalo III Formation of western Madagascar. Paleogeography During the Middle Jurassic Epoch, Pangaea began to separate into Laurasia and Gondwana, and the Atlantic Ocean formed. Eastern Laurasia was tectonically active as the Cimmerian plate continued to collide with Laurasia's southern coast, completely closing the Paleo-Tethys Ocean. A subduction zone ...
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Bathonian
In the geologic timescale the Bathonian is an age and stage of the Middle Jurassic. It lasted from approximately 168.3 Ma to around 166.1 Ma (million years ago). The Bathonian Age succeeds the Bajocian Age and precedes the Callovian Age. Stratigraphic definitions The Bathonian Stage takes its name from Bath, a spa town in England built on Jurassic limestone (the Latinized form of the town name is ''Bathonium''). The name was introduced in scientific literature by Belgian geologist d'Omalius d'Halloy in 1843. The original type locality was located near Bath. The French palaeontologist Alcide d'Orbigny was in 1852 the first to define the exact length of the stage. The base of the Bathonian is at the first appearance of ammonite species '' Parkinsonia (Gonolkites) convergens'' in the stratigraphic column. The global reference profile for the base of the Bathonian (a GSSP) was ratified as Ravin du Bès, Bas-Auran area, Alpes de Haute Provence, France in 2009. The top of the Bat ...
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Great Oolite Group
The Great Oolite Group is a Middle Jurassic stratigraphic unit that outcrops in southern England. It consists of a complex set of marine deposits primarily mudstone and bioclastic ooidal and fine grained limestone, deposited in nearshore to shelf settings. It is exposed at the surface as a variably thick belt extending roughly NE-SW from the coast of Dorset up to the Humber. It is also present at depth in the Weald and Wessex Basins, as well as offshore. Several of the constituent formations, notably the Taynton Limestone Formation and the Forest Marble Formation are notable for their fossil content, including those of dinosaurs and pterosaurs and some of the earliest mammals. Paleofauna * '' Cardiodon rugulosus'' * ''Cetiosaurus oxoniensis'' * '' Eoplophysis vetustus'' (stegosaurid indet) * ''Megalosaurus bucklandii'' * ''Proceratosaurus bradleyi'' * '' Seldsienean megistorhynchus'' See also * List of dinosaur-bearing rock formations ** List of stratigraphic units with fe ...
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Gloucestershire
Gloucestershire ( abbreviated Glos) is a county in South West England. The county comprises part of the Cotswold Hills, part of the flat fertile valley of the River Severn and the entire Forest of Dean. The county town is the city of Gloucester and other principal towns and villages include Cheltenham, Cirencester, Kingswood, Bradley Stoke, Stroud, Thornbury, Yate, Tewkesbury, Bishop's Cleeve, Churchdown, Brockworth, Winchcombe, Dursley, Cam, Berkeley, Wotton-under-Edge, Tetbury, Moreton-in-Marsh, Fairford, Lechlade, Northleach, Stow-on-the-Wold, Chipping Campden, Bourton-on-the-Water, Stonehouse, Nailsworth, Minchinhampton, Painswick, Winterbourne, Frampton Cotterell, Coleford, Cinderford, Lydney and Rodborough and Cainscross that are within Stroud's urban area. Gloucestershire borders Herefordshire to the north-west, Worcestershire to the north, Warwickshire to the north-east, Oxfordshire to the east, Wiltshire to the south, Bristol and Somerset ...
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Pterosaur
Pterosaurs (; from Greek ''pteron'' and ''sauros'', meaning "wing lizard") is an extinct clade of flying reptiles in the order, Pterosauria. They existed during most of the Mesozoic: from the Late Triassic to the end of the Cretaceous (228 to 66 million years ago). Pterosaurs are the earliest vertebrates known to have evolved powered flight. Their wings were formed by a membrane of skin, muscle, and other tissues stretching from the ankles to a dramatically lengthened fourth finger. There were two major types of pterosaurs. Basal pterosaurs (also called 'non-pterodactyloid pterosaurs' or 'rhamphorhynchoids') were smaller animals with fully toothed jaws and, typically, long tails. Their wide wing membranes probably included and connected the hind legs. On the ground, they would have had an awkward sprawling posture, but the anatomy of their joints and strong claws would have made them effective climbers, and some may have even lived in trees. Basal pterosaurs were insectiv ...
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Wastebasket Taxon
Wastebasket taxon (also called a wastebin taxon, dustbin taxon or catch-all taxon) is a term used by some taxonomists to refer to a taxon that has the sole purpose of classifying organisms that do not fit anywhere else. They are typically defined by either their designated members' often superficial similarity to each other, or their ''lack'' of one or more distinct character states or by their ''not'' belonging to one or more other taxa. Wastebasket taxa are by definition either paraphyletic or polyphyletic, and are therefore not considered valid taxa under strict cladistic rules of taxonomy. The name of a wastebasket taxon may in some cases be retained as the designation of an evolutionary grade, however. The term was coined in a 1985 essay by Steven Jay Gould. Examples There are many examples of paraphyletic groups, but true "wastebasket" taxa are those that are known not to, and perhaps not intended to, represent natural groups, but are nevertheless used as convenient groups ...
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Thalattosuchia
Thalattosuchia is a clade of marine crocodylomorphs from the Early Jurassic to the Early Cretaceous that had a cosmopolitan distribution. They are colloquially referred to as marine crocodiles or sea crocodiles, though they are not members of Crocodilia and records from Thailand and China suggest that some members lived in freshwater. The clade contains two major subgroupings, the Teleosauroidea and Metriorhynchoidea. Teleosauroids are not greatly specialised for oceanic life, with back osteoderms similar to other crocodyliformes. Within Metriorhynchoidea, the Metriorhynchidae displayed extreme adaptions for life in the open ocean, including the transformation of limbs into flippers, the development of a tail fluke, and smooth, scaleless skin. Discovery and naming The term Thalattosuchia was coined by Fraas in 1901.Fraas E. 1901. Die Meerkrokodile (Thalattosuchia n. g.) eine neue Sauriergruppe der Juraformation. ''Jahreshefte des Vereins für vaterländische Naturkunde, Württem ...
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Nomen Dubium
In binomial nomenclature, a ''nomen dubium'' (Latin for "doubtful name", plural ''nomina dubia'') is a scientific name that is of unknown or doubtful application. Zoology In case of a ''nomen dubium'' it may be impossible to determine whether a specimen belongs to that group or not. This may happen if the original type series (i. e. holotype, isotype, syntype or paratype) is lost or destroyed. The zoological and botanical codes allow for a new type specimen, or neotype, to be chosen in this case. A name may also be considered a ''nomen dubium'' if its name-bearing type is fragmentary or lacking important diagnostic features (this is often the case for species known only as fossils). To preserve stability of names, the ''International Code of Zoological Nomenclature'' allows a new type specimen, or neotype, to be chosen for a ''nomen dubium'' in this case. 75.5. Replacement of unidentifiable name-bearing type by a neotype. When an author considers that the taxonomic identity of a ...
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Middle Jurassic Reptiles Of Europe
Middle or The Middle may refer to: * Centre (geometry), the point equally distant from the outer limits. Places * Middle (sheading), a subdivision of the Isle of Man * Middle Bay (other) * Middle Brook (other) * Middle Creek (other) * Middle Island (other) * Middle Lake (other) * Middle Mountain, California * Middle Peninsula, Chesapeake Bay, Virginia * Middle Range, a former name of the Xueshan Range on Taiwan Island * Middle River (other) * Middle Rocks, two rocks at the eastern opening of the Straits of Singapore * Middle Sound, a bay in North Carolina * Middle Township (other) * Middle East Music * "Middle" (song), 2015 * "The Middle" (Jimmy Eat World song), 2001 * "The Middle" (Zedd, Maren Morris and Grey song), 2018 *"Middle", a song by Rocket from the Crypt from their 1995 album ''Scream, Dracula, Scream!'' *"The Middle", a song by Demi Lovato from their debut album ''Don't Forget'' *"The Middle", a song by T ...
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