Qianshanosuchus
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Qianshanosuchus
''Qianshanosuchus'' is a genus of basal crocodyloid from the Paleocene of the Qianshan Basin, China. The fossil material, which includes an incomplete skull and parts of the lower jaw, show various features usually associated with juvenile crocodiles alongside various unique traits that were used to erect a new genus. It is the first and only basal crocodyloid currently known from the Paleocene of China, which had previously only yielded alligatoroids and planocraniids. Its presence in this part of the world and its basal position to species of the genus ''Asiatosuchus'' supports the idea that crocodyloids dispersed from Asia into Europe. ''Qianshanosuchus'' only includes a single species, ''Qianshanosuchus youngi''. History and naming ''Qianshanosuchus'' is known from only a single specimen, IBCAS QS15, an incomplete skull and an associated part of the mandible. Based on the small size of the holotype and several anatomical traits, it is thought to preserve a juvenile specim ...
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Crocodyloidea
Crocodyloidea is one of three superfamilies of crocodilians, the other two being Alligatoroidea and Gavialoidea, and it includes the crocodiles. Crocodyloidea may also include the extinct Mekosuchinae, native to Australasia from the Eocene to the Holocene, although this is disputed. Classification Cladistically, it is defined as ''Crocodylus niloticus'' (the Nile crocodile) and all crocodylians more closely related to ''C. niloticus'' than to either ''Alligator mississippiensis'' (the American alligator) or ''Gavialis gangeticus'' (the gharial). This is a stem-based taxon, stem-based definition for crocodiles Crocodiles (family Crocodylidae) or true crocodiles are large semiaquatic reptiles that live throughout the tropics in Africa, Asia, the Americas and Australia. The term crocodile is sometimes used even more loosely to include all extant memb ..., and is more inclusive than the crown group Crocodylidae. As a crown group, Crocodylidae only includes the last common anc ...
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Crocodyloid
Crocodyloidea is one of three superfamilies of crocodilians, the other two being Alligatoroidea and Gavialoidea, and it includes the crocodiles. Crocodyloidea may also include the extinct Mekosuchinae, native to Australasia from the Eocene to the Holocene, although this is disputed. Classification Cladistically, it is defined as ''Crocodylus niloticus'' (the Nile crocodile) and all crocodylians more closely related to ''C. niloticus'' than to either ''Alligator mississippiensis'' (the American alligator) or ''Gavialis gangeticus'' (the gharial). This is a stem-based definition for crocodiles, and is more inclusive than the crown group Crocodylidae. As a crown group, Crocodylidae only includes the last common ancestor of all extant (living) crocodiles and their descendants (living or extinct), whereas Crocodyloidea, as a stem group, also includes more basal extinct crocodile ancestors that are more closely related to living crocodiles than to alligators or gavialids. When consi ...
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Gavialoidea
Gavialoidea is one of three superfamilies of crocodylians, the other two being Alligatoroidea and Crocodyloidea. Although many extinct species are known, only the gharial ''Gavialis gangeticus'' and the false gharial ''Tomistoma schlegelii'' are alive today, with ''Hanyusuchus'' having become extinct in the last few centuries. Extinct South American gavialoids likely dispersed in the mid Tertiary from Africa and Asia. Fossil remains of the Puerto Rican gavialoid '' Aktiogavialis puertorisensis'' were discovered in a cave located in San Sebastián, Puerto Rico and dated to the Oligocene. This individual is thought to have crossed the Atlantic coming from Africa, indicating that this species was able to withstand saltwater. Classification Gavialoidea is cladistically defined as ''Gavialis gangeticus'' (the gharial) and all crocodylians closer to it than to ''Alligator mississippiensis'' (the American alligator) or ''Crocodylus niloticus'' (the Nile crocodile). This is a stem-bas ...
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Crocodylinae
Crocodylinae is a subfamily of true crocodiles within the family Crocodylidae, and is the sister taxon to Osteolaeminae ( dwarf crocodiles and slender-snouted crocodiles). Taxonomy Crocodylinae was cladistically defined by Christopher Brochu in 1999 as ''Crocodylus niloticus'' (the Nile crocodile) and all crocodylians more closely related to it than to ''Osteolaemus tetraspis'' (the Dwarf crocodile). This is a stem-based definition, and is the sister taxon to Osteolaeminae. Crocodylinae contains the extant genus '' Crocodylus''. It is disputed as to whether is also includes ''Mecistops'' (slender-snouted crocodiles), or the extinct genus Voay. Phylogeny Some morphological studies have recovered ''Mecistops'' as a basal member of Crocodylinae, more closely related to '' Crocodylus'' than to ''Osteolaemus'' and the other members of Osteolaeminae, as shown in the cladogram below. The below cladogram is based on a 2021 study using paleogenomics that extracted DNA from the ext ...
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Osteolaeminae
Osteolaeminae is a subfamily of true crocodiles within the family Crocodylidae containing the dwarf crocodiles and slender-snouted crocodiles, and is the sister taxon to Crocodylinae. Taxonomy Osteolaeminae was named by Christopher Brochu in 2003 as a subfamily of Crocodylidae separate from Crocodylinae, and is cladistically defined as ''Osteolaemus tetraspis'' (the Dwarf crocodile) and all crocodylians more closely related to it than to ''Crocodylus niloticus'' (the Nile crocodile). This is a stem-based definition, and is the sister taxon to Crocodylinae. Osteolaeminae contains the two extant genera ''Osteolaemus'' and ''Mecistops'', along with several extinct genera, although the number of extant species within Osteolaeminae is currently in question. Phylogeny The cladogram below is based on two studies that combined morphological, molecular (DNA sequencing), and stratigraphic (fossil age) data. Alternatively, other morphological studies have recovered ''Mecistops'' a ...
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Mekosuchinae
Mekosuchinae is an extinct clade of crocodilians from the Cenozoic of Australasia. They first appear in the fossil record in the Eocene in Australia, and survived until the arrival of humans: in the Pleistocene in Australia and within the Holocene in the Pacific islands of Fiji, New Caledonia and Vanuatu. Mekosuchine crocodiles are a diverse group. One of the last species, ''Mekosuchus inexpectatus'' from Holocene New Caledonia, may have been arboreal. The early Miocene species '' Harpacochampsa camfieldensis'' may have resembled a false gharial. Another mekosuchine fossil, currently undescribed, has been found in Miocene deposits from New Zealand. One genus, ''Mekosuchus'', managed to spread to the islands of the Pacific; it is believed to have island-hopped across the Coral Sea, moving first to a now submerged island known as Greater Chesterfield Island, then New Caledonia and onwards. In the Pleistocene, ''Quinkana'' was one of the top terrestrial predators of the Australian ...
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Asiatosuchus Grangeri
''Asiatosuchus'' is an extinct genus of crocodyloid crocodilian Crocodilia (or Crocodylia, both ) is an order of mostly large, predatory, semiaquatic reptiles, known as crocodilians. They first appeared 95 million years ago in the Late Cretaceous period ( Cenomanian stage) and are the closest living ...s that lived in Eurasia during the Paleogene. Many Paleogene crocodilians from Europe and Asia have been attributed to ''Asiatosuchus'' since the genus was named in 1940. These species have a generalized crocodilian morphology (biology), morphology typified by flat, triangular skulls. The feature that traditionally united these species under the genus ''Asiatosuchus'' is a broad connection or Symphysis menti, symphysis between the two halves of the lower jaw. Recent studies of the evolutionary relationships of early crocodilians along with closer examinations of the morphology of fossil specimens suggest that only the first named species of ''Asiatosuchus'', ''A. grange ...
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