Protuberum
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Protuberum
''Protuberum'' is an extinct genus of traversodontid cynodonts known from a single species ''Protuberum cabralense'',The specific name ''cabralense'' is "necessarily emended from the original ''P. cabralensis'' because ''Protuberum'' is a neuter generic name". from the Middle Triassic of Brazil., , & 2009 “A New Traversodontid Cynodont (Therapsida, Eucynodontia) from the Middle Triassic Santa Maria Formation of Rio Grande do Sul, Brazil” ''Palaeontology'' 52(1):229-250 As with all genera of the family Traversodontidae, ''Protuberum'' was a herbivore, with a specialized grinding action when feeding. The two known specimens were collected a number of years apart from sediments of the Alemoa Member of the Santa Maria Formation in Geopark of Paleorrota, Rio Grande do Sul, Brazil. The first specimen was collected in 1977 and consists of several ribs and vertebrae. The second specimen, collected in 1989, consists of a partial articulated skeleton and a skull. Both specimens ...
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2009 In Paleontology
Arthropods Cephalopods Three new species of extinct octopus, Octopoda discovered in 2009. The species – ''Keuppia hyperbolaris'', ''Keuppia levante'', and ''Styletoctopus annae'' – lived about 95 million years ago, and bear a strong resemblance to modern octopuses, suggesting that the Octopoda order has remained relatively unchanged for tens of millions of years. The fossils included evidence of cephalopod arm, arms, muscles, rows of suckers, cephalopod ink, ink, and internal gills. The discovery was made by a team led by Dirk Fuchs of the Freie University, which is located at Berlin, Germany.Rare fossil octopuses found
NBC News, March 18, 2009
The fossils were found at Hakel and Hadjoula, Lebanon.
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Cynodont
The cynodonts () (clade Cynodontia) are a clade of eutheriodont therapsids that first appeared in the Late Permian (approximately 260 mya), and extensively diversified after the Permian–Triassic extinction event. Cynodonts had a wide variety of lifestyles, including carnivory and herbivory. Mammals are cynodonts, as are their extinct ancestors and close relatives, having evolved from advanced probainognathian cynodonts during the Late Triassic. All other cynodont lines went extinct, with the last known non-mammalian cynodont group, the Tritylodontidae, having its youngest records in the Early Cretaceous. Description Early cynodonts have many of the skeletal characteristics of mammals. The teeth were fully differentiated and the braincase bulged at the back of the head. Outside of some crown-group mammals (notably the therians), all cynodonts probably laid eggs. The temporal fenestrae were much larger than those of their ancestors, and the widening of the zygomatic arch in ...
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Traversodontids
Traversodontidae is an extinct family of herbivorous cynodonts. Traversodonts were primarily Gondwanan, with many species known from Africa and South America. Recently, traversodonts have also been found from Europe and eastern North America. Traversodonts first appeared in the Middle Triassic and diversified in the Late Triassic before going extinct at the end of the epoch. The family Traversodontidae was erected by Friedrich von Huene in 1936 for cynodonts first found in São Pedro do Sul in Paleorrota, Brazil. Description Traversodonts are members of Gomphodontia, a group of herbivorous cynognathian cynodonts. As an adaptation toward eating plants, they have wide postcanine teeth behind large canines. These postcanines are closely spaced with their crowns touching each other. Each is usually wider than it is long and is covered in several cusps. Because of their complexity, postcanine teeth are the primary means of identifying and distinguishing different species of tra ...
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Santa Maria Formation
The Santa Maria Formation is a sedimentary rock formation found in Rio Grande do Sul, Brazil. It is primarily Carnian in age (Late Triassic), and is notable for its fossils of cynodonts, "rauisuchian" pseudosuchians, and early dinosaurs and other dinosauromorphs, including the herrerasaurid '' Staurikosaurus'', the basal sauropodomorphs ''Buriolestes'' and '' Saturnalia,'' and the lagerpetid ''Ixalerpeton''. The formation is named after the city of Santa Maria in the central region of Rio Grande do Sul, where outcrops were first studied. The Santa Maria Formation makes up the majority of the Santa Maria Supersequence, which extends through the entire Late Triassic. The Santa Maria Supersequence is divided into four geological sequences, separated from each other by short unconformities. The first two of these sequences (Pinheiros-Chiniquá and Santa Cruz sequences) lie entirely within the Santa Maria Formation, while the third (the Candelária sequence) is shared with the ov ...
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Traversodontidae
Traversodontidae is an extinct family of herbivorous cynodonts. Traversodonts were primarily Gondwanan, with many species known from Africa and South America. Recently, traversodonts have also been found from Europe and eastern North America. Traversodonts first appeared in the Middle Triassic and diversified in the Late Triassic before going extinct at the end of the epoch. The family Traversodontidae was erected by Friedrich von Huene in 1936 for cynodonts first found in São Pedro do Sul in Paleorrota, Brazil. Description Traversodonts are members of Gomphodontia, a group of herbivorous cynognathian cynodonts. As an adaptation toward eating plants, they have wide postcanine teeth behind large canines. These postcanines are closely spaced with their crowns touching each other. Each is usually wider than it is long and is covered in several cusps. Because of their complexity, postcanine teeth are the primary means of identifying and distinguishing different species of traverso ...
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Middle Triassic
In the geologic timescale, the Middle Triassic is the second of three epochs of the Triassic period or the middle of three series in which the Triassic system is divided in chronostratigraphy. The Middle Triassic spans the time between Ma and Ma (million years ago). It is preceded by the Early Triassic Epoch and followed by the Late Triassic Epoch. The Middle Triassic is divided into the Anisian and Ladinian ages or stages. Formerly the middle series in the Triassic was also known as Muschelkalk. This name is now only used for a specific unit of rock strata with approximately Middle Triassic age, found in western Europe. Middle Triassic fauna Following the Permian–Triassic extinction event, the most devastating of all mass-extinctions, life recovered slowly. In the Middle Triassic, many groups of organisms reached higher diversity again, such as the marine reptiles (e.g. ichthyosaurs, sauropterygians, thallatosaurs), ray-finned fish and many invertebrate groups like ...
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Brazil
Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area and the seventh most populous. Its capital is Brasília, and its most populous city is São Paulo. The federation is composed of the union of the 26 States of Brazil, states and the Federal District (Brazil), Federal District. It is the largest country to have Portuguese language, Portuguese as an List of territorial entities where Portuguese is an official language, official language and the only one in the Americas; one of the most Multiculturalism, multicultural and ethnically diverse nations, due to over a century of mass Immigration to Brazil, immigration from around the world; and the most populous Catholic Church by country, Roman Catholic-majority country. Bounded by the Atlantic Ocean on the east, Brazil has a Coastline of Brazi ...
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Herbivore
A herbivore is an animal anatomically and physiologically adapted to eating plant material, for example foliage or marine algae, for the main component of its diet. As a result of their plant diet, herbivorous animals typically have mouthparts adapted to rasping or grinding. Horses and other herbivores have wide flat teeth that are adapted to grinding grass, tree bark, and other tough plant material. A large percentage of herbivores have mutualistic gut flora that help them digest plant matter, which is more difficult to digest than animal prey. This flora is made up of cellulose-digesting protozoans or bacteria. Etymology Herbivore is the anglicized form of a modern Latin coinage, ''herbivora'', cited in Charles Lyell's 1830 ''Principles of Geology''.J.A. Simpson and E.S.C. Weiner, eds. (2000) ''The Oxford English Dictionary'', vol. 8, p. 155. Richard Owen employed the anglicized term in an 1854 work on fossil teeth and skeletons. ''Herbivora'' is derived from Latin ''herba' ...
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Type Species
In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen(s). Article 67.1 A similar concept is used for suprageneric groups and called a type genus. In botanical nomenclature, these terms have no formal standing under the code of nomenclature, but are sometimes borrowed from zoological nomenclature. In botany, the type of a genus name is a specimen (or, rarely, an illustration) which is also the type of a species name. The species name that has that type can also be referred to as the type of the genus name. Names of genus and family ranks, the various subdivisions of those ranks, and some higher-rank names based on genus names, have such types.
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Geopark
A geopark is a protected area with internationally significant geology within which sustainable development is sought and which includes tourism, conservation, education and research concerning not just geology but other relevant sciences. In 2005, a European Geopark was defined as being: "a territory with a particular geological heritage and with a sustainable territorial development....the ultimate aim of a European Geopark is to bring enhanced employment opportunities for the people who live there." Today the geopark is virtually synonymous with the UNESCO geopark, which is defined and managed under the voluntary authority of UNESCO's International Geoscience and Geoparks Programme (IGGP). UNESCO provides a standard for geoparks and a certification service to parks that apply for it. The service is available to member states of UNESCO. This list is not the same as the member states of the United Nations. Membership in the UN does not automatically imply membership in UNESCO ...
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Rio Grande Do Sul
Rio Grande do Sul (, , ; "Great River of the South") is a Federative units of Brazil, state in the South Region, Brazil, southern region of Brazil. It is the Federative_units_of_Brazil#List, fifth-most-populous state and the List of Brazilian states by area, ninth largest by area. Located in the southernmost part of the country, Rio Grande do Sul is bordered clockwise by Santa Catarina (state), Santa Catarina to the north and northeast, the Atlantic Ocean to the east, the Uruguayan Departments of Uruguay, departments of Rocha Department, Rocha, Treinta y Tres Department, Treinta y Tres, Cerro Largo Department, Cerro Largo, Rivera Department, Rivera and Artigas Department, Artigas to the south and southwest, and the Argentina, Argentine Provinces of Argentina, provinces of Corrientes Province, Corrientes and Misiones Province, Misiones to the west and northwest. The capital and largest city is Porto Alegre. The state has the highest life expectancy in Brazil, and the crime rate i ...
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