Progymnosperms
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Progymnosperms
The progymnosperms are an extinct group of woody, spore-bearing plants that is presumed to have evolved from the trimerophytes, and eventually gave rise to the gymnosperms, ancestral to acrogymnosperms and angiosperms (flowering plants). They have been treated formally at the rank of division Progymnospermophyta or class Progymnospermopsida (as opposite). The stratigraphically oldest known examples belong to the Middle Devonian order the Aneurophytales, with forms such as '' Protopteridium'', in which the vegetative organs consisted of relatively loose clusters of axes. ''Tetraxylopteris'' is another example of a genus lacking leaves. In more advanced aneurophytaleans such as ''Aneurophyton'' these vegetative organs started to look rather more like fronds, and eventually during Late Devonian times the aneurophytaleans are presumed to have given rise to the pteridosperm order, the Lyginopteridales. In Late Devonian times, another group of progymnosperms gave rise to the first re ...
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Archaeopteris
''Archaeopteris'' is an extinct genus of progymnosperm tree with fern-like leaves. A useful index fossil, this tree is found in strata dating from the Upper Devonian to Lower Carboniferous (), the oldest fossils being 385 million years old, and had global distribution. Until the 2007 discovery of '' Wattieza'', many scientists considered ''Archaeopteris'' to be the earliest known tree. Bearing buds, reinforced branch joints, and branched trunks similar to today's wood, it is more reminiscent of modern seed-bearing trees than other spore bearing taxa; It combines characteristics of woody trees and herbaceous ferns, and belongs to the progymnosperms, a group of extinct plants with gymnosperm-like wood but that produce spores rather than seeds. Taxonomy John William Dawson described the genus in 1871. The name derives from the ancient Greek (''archaīos'', "ancient"), and (''ptéris'', "fern"). ''Archaeopteris'' was originally classified as a fern, and it remained classif ...
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Heterosporous
Heterospory is the production of spores of two different sizes and sexes by the sporophytes of land plants. The smaller of these, the microspore, is male and the larger megaspore is female. Heterospory evolved during the Devonian period from isospory independently in several plant groups: the clubmosses, the ferns including the arborescent horsetails, and progymnosperms. This occurred as part of the process of evolution of the timing of sex differentiation.Sussex, I.M. (1966) The origin and development of heterospory in vascular plants. Chapter 9 in ''Trends in Plant morphogenesis'', ed. by E.G. Cutter, Longmans. Origin of heterospory Heterospory evolved due to natural selection that favoured an increase in propagule size compared with the smaller spores of homosporous plants. Heterosporous plants, similar to anisosporic plants, produce two different sized spores in separate sporangia that develop into separate male and female gametophytes. It is proposed that the emergence ...
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Lyginopteridales
The Lyginopteridales were the archetypal pteridosperms: They were the first plant fossils to be described as pteridosperms and, thus, the group on which the concept of pteridosperms was first developed;Oliver, F. W. & Scott, D. H. (1904). "On the structure of the Palaeozoic seed ''Lagenostoma Lomaxi'', with a statement of the evidence upon which it is referred to ''Lyginodendron''." ''Philosophical Transactions of the Royal Society of London, Series B,'' 197: 193-247. they are the stratigraphically oldest-known pteridosperms, occurring first in late Devonian strata;Rothwell, G. W., Scheckler, S. E. & Gillespie, W. H. (1989). "''Elkinsia'' gen. nov., a Late Devonian gymnosperm with cupulate ovules." ''Botanical Gazette,'' 150: 170-189. and they have the most primitive features, most notably in the structure of their ovules.Long, A. G. (1959). "On the structure of ''Calymmatotheca kidstoni'' Calder (emended) and ''Genomosperma latens'' gen. et sp. nov. from the Calciferous Sand ...
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Gymnosperms
The gymnosperms ( lit. revealed seeds) are a group of seed-producing plants that includes conifers, cycads, ''Ginkgo'', and gnetophytes, forming the clade Gymnospermae. The term ''gymnosperm'' comes from the composite word in el, γυμνόσπερμος ( el, γυμνός, translit=gymnos, lit=naked, label=none and el, σπέρμα, translit=sperma, lit=seed, label=none), literally meaning 'naked seeds'. The name is based on the unenclosed condition of their seeds (called ovules in their unfertilized state). The non-encased condition of their seeds contrasts with the seeds and ovules of flowering plants (angiosperms), which are enclosed within an ovary. Gymnosperm seeds develop either on the surface of scales or leaves, which are often modified to form cones, or solitary as in yew, ''Torreya'', ''Ginkgo''. Gymnosperm lifecycles involve alternation of generations. They have a dominant diploid sporophyte phase and a reduced haploid gametophyte phase which is dependent on ...
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Tetraxylopteris
'' Tetraxylopteris'' is a genus of extinct vascular plants of the Middle to Upper Devonian (around ). Fossils were first found in New York State, USA. A second species was later found in Venezuela. Description Fossils of ''Tetraxylopteris'' have so far been discovered in two locations. ''T. schmidtii'' was named from the Catskill Clastic Wedge, New York State, United States of America, in rocks of Middle to Upper Devonian age (around ). ''T. reposana'' was found in the Campo Chico Formation, north-west Venezuela, in beds believed to be of Frasnian age (). The overall shape of ''Tetraxylopteris'' consisted of a complex system of branches. The main stem was 'pseudomonopodial', i.e. it divided dichotomously to produce side stems while the main stem maintained its identity. The main and side stems then bore three orders of branches (i.e. the first branches from the stems divided twice more). (There were possibly four orders in ''T. reposana''.) The branches were arranged in opposit ...
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Monopodial
Vascular plants with monopodial growth habits grow upward from a single point. They add leaves to the apex each year and the stem grows longer accordingly. The word ''Monopodial'' is derived from Greek "mono-", ''one'' and "podial", "foot", in reference to the fact that monopodial plants have a single trunk or stem. Orchids with monopodial growth often produce copious aerial roots that often hang down in long drapes and have green chlorophyll underneath the grey root coverings, which are used as additional photosynthetic organs. They do not have a rhizome or pseudobulbs so species adapted to dry periods have fleshy succulent leaves instead. Flowers generally come from the stem between the leaves. With some monopodial species, the stem (the rhizome) might fork into two, but for all monopodial orchids this is not necessary for continued growth, as opposed to orchids with sympodial growth.''The Orchid Expert'', Dr. D. G. Hessayon David Gerald Hessayon OBE (born 1928) is a British a ...
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Conifer
Conifers are a group of conifer cone, cone-bearing Spermatophyte, seed plants, a subset of gymnosperms. Scientifically, they make up the phylum, division Pinophyta (), also known as Coniferophyta () or Coniferae. The division contains a single extant class (biology), class, Pinopsida. All Neontology, extant conifers are perennial plant, perennial woody plants with secondary growth. The great majority are trees, though a few are shrubs. Examples include Cedrus, cedars, Pseudotsuga, Douglas-firs, Cupressaceae, cypresses, firs, junipers, Agathis, kauri, larches, pines, Tsuga, hemlocks, Sequoioideae, redwoods, spruces, and Taxaceae, yews.Campbell, Reece, "Phylum Coniferophyta". Biology. 7th. 2005. Print. P. 595 As of 1998, the division Pinophyta was estimated to contain eight families, 68 genera, and 629 living species. Although the total number of species is relatively small, conifers are ecology, ecologically important. They are the dominant plants over large areas of land, most ...
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Spruce
A spruce is a tree of the genus ''Picea'' (), a genus of about 35 species of coniferous evergreen trees in the family Pinaceae, found in the northern temperate and boreal (taiga) regions of the Earth. ''Picea'' is the sole genus in the subfamily Piceoideae. Spruces are large trees, from about 20 to 60 m (about 60–200 ft) tall when mature, and have whorled branches and conical form. They can be distinguished from other members of the pine family by their needles (leaves), which are four-sided and attached singly to small persistent peg-like structures (pulvini or sterigmata) on the branches, and by their cones (without any protruding bracts), which hang downwards after they are pollinated. The needles are shed when 4–10 years old, leaving the branches rough with the retained pegs. In other similar genera, the branches are fairly smooth. Spruce are used as food plants by the larvae of some Lepidoptera (moth and butterfly) species, such as the eastern spruce budwo ...
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Macrophyll
Leaf size of plants can be described using the terms ''megaphyll'', ''macrophyll'', ''mesophyll'', ''microphyll'', ''nanophyll'' and ''leptophyll'' (in descending order) in a classification devised in 1934 by Christen C. Raunkiær and since modified by others. Definitions vary, some referring to length and others to area. Raunkiaer's original definitions were by leaf area, and differed by a factor of nine at each stage. Some authors simplified the system to make it specific to particular climates, and have introduced extra terms including notophyll, ''picophyll'', ''platyphyll'' and ''subleptophyll''. In ecology, microphyll and similar terms based on blade size of the leaf are used to describe a flora, for example, a "microphyll rainforest" is often defined as a forest where the dominant trees have leaves less than 7.5 cm in length. Raunkiaer's work Christen C. Raunkiaer proposed using leaf size as a relatively easy measurement that could be used to compare the adaptation of ...
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Phloem
Phloem (, ) is the living biological tissue, tissue in vascular plants that transports the soluble organic compounds made during photosynthesis and known as ''photosynthates'', in particular the sugar sucrose, to the rest of the plant. This transport process is called translocation. In trees, the phloem is the innermost layer of the bark (botany), bark, hence the name, derived from the Ancient Greek word (''phloiós''), meaning "bark". The term was introduced by Carl Nägeli in 1858. Structure Phloem tissue consists of conducting Cell (biology), cells, generally called sieve elements, Ground tissue#Parenchyma, parenchyma cells, including both specialized companion cells or albuminous cells and unspecialized cells and supportive cells, such as fibres and sclereids. Conducting cells (sieve elements) Sieve elements are the type of cell that are responsible for transporting sugars throughout the plant. At maturity they lack a Cell nucleus, nucleus and have very few organelles, ...
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Homosporous
In biology, a spore is a unit of sexual or asexual reproduction that may be adapted for dispersal and for survival, often for extended periods of time, in unfavourable conditions. Spores form part of the life cycles of many plants, algae, fungi and protozoa. Bacterial spores are not part of a sexual cycle, but are resistant structures used for survival under unfavourable conditions. Myxozoan spores release amoeboid infectious germs ("amoebulae") into their hosts for parasitic infection, but also reproduce within the hosts through the pairing of two nuclei within the plasmodium, which develops from the amoebula. In plants, spores are usually haploid and unicellular and are produced by meiosis in the sporangium of a diploid sporophyte. Under favourable conditions the spore can develop into a new organism using mitotic division, producing a multicellular gametophyte, which eventually goes on to produce gametes. Two gametes fuse to form a zygote which develops into a ne ...
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