Poposauroidea
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Poposauroidea
Poposauroidea is a clade of advanced pseudosuchians (archosaurs closer to crocodilians than to dinosaurs). It includes poposaurids, shuvosaurids, ctenosauriscids, and other unusual pseudosuchians such as ''Qianosuchus'' and ''Lotosaurus''. However, it excludes most large predatory quadrupedal "rauisuchians" such as rauisuchids and "prestosuchids". Those reptiles are now allied with crocodylomorphs (crocodile ancestors) in a clade known as Loricata, which is the sister taxon to the poposauroids in the clade Paracrocodylomorpha. Although it was first formally defined in 2007, the name "Poposauroidea" has been used for many years. The group has been referred to as Poposauridae by some authors, although this name is often used more narrowly to refer to the family that includes ''Poposaurus'' and its close relatives. Poposauroids went extinct at the end of the Triassic period along with other non-crocodylomorph pseudosuchians. However, they were among the most diverse and long-lasting ...
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Rauisuchia
"Rauisuchia" is a paraphyletic group of mostly large and carnivorous Triassic archosaurs. Rauisuchians are a category of archosaurs within a larger group called Pseudosuchia, which encompasses all archosaurs more closely related to crocodilia Crocodilia (or Crocodylia, both ) is an order of mostly large, predatory, semiaquatic reptiles, known as crocodilians. They first appeared 95 million years ago in the Late Cretaceous period ( Cenomanian stage) and are the closest living ...ns than to birds and other dinosaurs. First named in the 1940s, Rauisuchia was a name exclusive to Triassic archosaurs which were generally large (often ), carnivorous, and quadrupedal with a pillar-erect hip posture, though exceptions exist for all of these traits. Rauisuchians, as a traditional Taxonomy, taxonomic group, were considered distinct from other Triassic archosaur groups such as early dinosaurs, Phytosaur, phytosaurs (crocodile-like carnivores), Aetosaur, aetosaurs (armored herbivor ...
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Poposaurus
''Poposaurus'' (" Popo Agie reptile") is an extinct genus of pseudosuchian archosaur from the Late Triassic of the southwestern United States. It belongs to the clade Poposauroidea, an unusual group of Triassic pseudosuchians that includes sail-backed, beaked, and aquatic forms. Fossils have been found in Wyoming, Utah, Arizona, and Texas. Except for the skull, most parts of the skeleton are known. The type species, ''P. gracilis'', was described and named by Maurice Goldsmith Mehl in 1915. A second species, ''P. langstoni'', was originally the type species of the genus ''Lythrosuchus''. Since it was first described, ''Poposaurus'' has been variously classified as a dinosaur, a phytosaur, and a "rauisuchian". Like theropod dinosaurs, ''Poposaurus'' was an obligate biped, meaning that it walked on two legs rather than four. However, as a pseudosuchian, it is more closely related to living crocodilians than to dinosaurs. ''Poposaurus'' is thought to have evolved this form of locomo ...
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Lotosaurus
''Lotosaurus'' is an extinct genus of sail-backed poposauroid Poposauroidea is a clade of advanced pseudosuchians (archosaurs closer to crocodilians than to dinosaurs). It includes poposaurids, shuvosaurids, ctenosauriscids, and other unusual pseudosuchians such as ''Qianosuchus'' and ''Lotosaurus''. Howeve ... known from Hunan Province of central China. Discovery ''Lotosaurus'' is known from the holotype IVPP V 4881 (or possibly V 4880), an articulated and well-preserved skeleton. Other referred specimens include IVPP V 48013 (a skull) as well as many articulated and disarticulated skeletal remains concentrated in a bonebed which is almost completely composed of ''Lotosaurus'' bones. All known specimens of this genus were collected from this bonebed, known as the ''Lotosaurus'' site, which belongs to the Batung Formation (or alternatively Xinlingzhen Formation of the Badong Group). Further excavations in 2018 revealed many more specimens, as well as ...
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Paracrocodylomorpha
Paracrocodylomorpha is a clade of pseudosuchian archosaurs. The clade includes the diverse and unusual group Poposauroidea as well as the generally carnivorous and quadrupedal members of Loricata, including modern crocodylians. Paracrocodylomorpha was named by paleontologist J. Michael Parrish in 1993, although the group is now considered to encompass more reptiles than his original definition intended. The most recent definition of Paracrocodylomorpha, as defined by Sterling Nesbitt in 2011, is "the least inclusive clade containing ''Poposaurus'' and ''Crocodylus niloticus'' (the Nile crocodile). Most groups of paracrocodylomorphs became extinct at the end of the Triassic period, with the exception of the crocodylomorphs, from which crocodylians such as crocodiles and alligators evolved in the latter part of the Mesozoic. History and definition Parrish (1993) defined Paracrocodylomorpha as the last common ancestor of "Poposauridae" and Crocodylomorpha and all of its descendants ...
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Loricata
Loricata is a clade of archosaur reptiles that includes crocodilians and some of their Triassic relatives, such as ''Postosuchus'' and ''Prestosuchus''. More specifically, Loricata includes Crocodylomorpha (the persistent archosaur subset which crocodilians belong to) and most "rauisuchians", a paraphyletic grade of large terrestrial pseudosuchians which were alive in the Triassic period and ancestral to crocodylomorphs. Loricata is one branch of the group Paracrocodylomorpha; the other branch is the clade Poposauroidea, an unusual collection of strange "rauisuchians" including bipedal, herbivorous, and sail-backed forms. The vast majority of typical "rauisuchians", which were usually quadrupedal predators, occupy basal (early-branching) rungs of Loricata leading up to crocodylomorphs. History and Classification Loricata was initially named in a completely different context by German naturalist Blasius Merrem in his 1820 ''Versuch eines Systems der Amphibien''. Merrem consid ...
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Ctenosauriscidae
Ctenosauriscidae is an extinct family (biology), family of pseudosuchian archosaurs within the clade Poposauroidea. Ctenosauriscids existed in Africa, Asia, Europe and North America during the Early Triassic to the Middle Triassic period (latest Olenekian to Anisian stages). All species had large "sails" on their backs. Ctenosauriscids are among some of the earliest archosaurs and represent the first global radiation of the group. Genera Phylogeny Ctenosauriscidae was named by Oskar Kuhn in 1964 in paleontology, 1964 to include the genus ''Ctenosauriscus''. It is a stem-based taxon defined by Richard J. Butler, Stephen L. Brusatte, Mike Reich, Sterling J. Nesbitt, Rainer R. Schoch and Jahn J. Hornung in 2011 as "the most inclusive clade containing ''Ctenosauriscus koeneni'' but not ''Poposaurus gracilis'', ''Effigia okeeffeae'', ''Postosuchus kirkpatricki'', ''Crocodylus niloticus'', ''Ornithosuchus longidens'', or ''Aetosaurus ferratus''". The cladograms below follows a 2011 ...
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Ctenosauriscid
Ctenosauriscidae is an extinct family (biology), family of pseudosuchian archosaurs within the clade Poposauroidea. Ctenosauriscids existed in Africa, Asia, Europe and North America during the Early Triassic to the Middle Triassic period (latest Olenekian to Anisian stages). All species had large "sails" on their backs. Ctenosauriscids are among some of the earliest archosaurs and represent the first global radiation of the group. Genera Phylogeny Ctenosauriscidae was named by Oskar Kuhn in 1964 in paleontology, 1964 to include the genus ''Ctenosauriscus''. It is a stem-based taxon defined by Richard J. Butler, Stephen L. Brusatte, Mike Reich, Sterling J. Nesbitt, Rainer R. Schoch and Jahn J. Hornung in 2011 as "the most inclusive clade containing ''Ctenosauriscus koeneni'' but not ''Poposaurus gracilis'', ''Effigia okeeffeae'', ''Postosuchus kirkpatricki'', ''Crocodylus niloticus'', ''Ornithosuchus longidens'', or ''Aetosaurus ferratus''". The cladograms below follows a 2011 ...
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Effigia
''Effigia'' was an extinct genus of shuvosaurid known from the Late Triassic of New Mexico, south-western USA. With a bipedal stance, long neck, and a toothless beaked skull, ''Effigia'' and other shuvosaurids bore a resemblance to the ornithomimid dinosaurs of the Cretaceous Period. However, shuvosaurids were not dinosaurs, but were instead a specialized family of poposauroid pseudosuchians, meaning that their closest living relatives are crocodilians. Discovery The 2-meter (6 ft 7 in) holotype fossil was collected by Edwin H. Colbert in 1947. At this time Colbert led an excavation to collect blocks of rock from the Whitaker Quarry of Ghost Ranch, near Abiquiu, New Mexico. Colbert's expedition intended to recover abundant fossils of the basal theropod dinosaur ''Coelophysis'', and he believed that no other large vertebrates were present in the quarry. As a result, his crew did not even open the plaster jackets of most of the blocks that were shipped to the American Museum of ...
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Xilousuchus
''Xilousuchus'' is an extinct genus of poposauroid from lower Triassic (Olenekian stage) deposits of Fugu County of northeastern Shanxi Province, China. It is known from the holotype, IVPP V 6026, a single well-preserved partial skeleton including the skull. It was found from the Heshanggou Formation of the Ordos Basin, Hazhen commune. It was first named by Xiao-Chun Wu in 1981 and the type species is ''Xilousuchus sapingensis''. Wu (1981) referred ''Xilousuchus'' to the Proterosuchia. Gower and Sennikov (1996) found it to be an erythrosuchian based strictly on the braincase. A more detailed re-description of the genus was provided by Nesbitt ''et al.'' (2010) and found poposauroid affinities. In his massive revision of archosaurs which included a large cladistic analysis, Sterling J. Nesbitt (2011) found ''Xilousuchus'' to be a poposauroid which is most closely related to ''Arizonasaurus''. ''Xilousuchus'' is the oldest archosaur to date, although ''Ctenosauriscus'' ...
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Pseudosuchia
Pseudosuchia is one of two major divisions of Archosauria, including living crocodilians and all archosaurs more closely related to crocodilians than to birds. Pseudosuchians are also informally known as "crocodilian-line archosaurs". Prior to 2011, the clade Pseudosuchia was often called Crurotarsi in reference to the crurotarsal ankle found in almost all members of the group, which traditionally included phytosaurs, ornithosuchids, and suchians. However, a major 2011 study of Triassic archosaur relations proposed that phytosaurs were not closely related to other traditional "crurotarsans", at least compared to "bird-line archosaurs" (Avemetatarsalians) such as pterosaurs and dinosaurs. As a result, the possession of a crurotarsal ankle was considered a plesiomorphic ("primitive") feature retained by pseudosuchians. Crurotarsi now refers to a broader group of reptiles including Pseudosuchia, Phytosauria, and Avemetatarsalia. Despite Pseudosuchia meaning "false crocodiles", th ...
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Qianosuchus
''Qianosuchus'' is an extinct genus of aquatic poposauroid archosaur from the middle Triassic (Anisian) Guanling Formation of Pan County, China. It is represented by two nearly complete skeletons and a crushed skull preserved in the limestone. ''Qianosuchus'' was at least 3 metres long, and had several skeletal adaptations which indicate a semi-marine lifestyle, similar to modern-day saltwater crocodiles. These adaptations have not been seen in any other archosaur from the Triassic. Description ''Qianosuchus'' had a skull around 33 cm (13 inches) long, with an elongated snout. The rostrum formed by the premaxilla is shallow at the front of the skull but deepens posteriorly. Each premaxilla has nine long teeth, and the maxillae bear eighteen teeth each. All the teeth are laterally compressed, curved backwards and serrated, like those of most other carnivorous archosaurs. The nares are expanded and elongated and almost collide with the antorbital fenestrae, meaning that the ...
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Shuvosauridae
Shuvosauridae is an extinct family of theropod-like pseudosuchians within the clade Poposauroidea. Shuvosaurids existed in North America (United States) and South America (Argentina) during the Late Triassic period (late Carnian to Rhaetian stages). Shuvosauridae was named by Sankar Chatterjee in 1993 to include the genus ''Shuvosaurus''. In a 2007 study ''Chatterjeea'' was demonstrated to be a junior synonym of ''Shuvosaurus'', and the therein cladistic analysis found that ''Shuvosaurus'', ''Effigia'' and ''Sillosuchus'' all form a closely related group. However, this group was left nameless, and simply referred to as Group Y. However, in accordance with ICZN rules of naming priority Shuvosauridae has priority over Chatterjeeidae (Shuvosauridae being named in 1993, while Chatterjeeidae was named in 1995). In 2011, Sterling J. Nesbitt proposed a new definition to this clade: "The least inclusive clade containing ''Shuvosaurus inexpectatus'' Chatterjee, 1993, and ''Sillosuchus ' ...
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