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Ctenosauriscidae
Ctenosauriscidae is an extinct family (biology), family of pseudosuchian archosaurs within the clade Poposauroidea. Ctenosauriscids existed in Africa, Asia, Europe and North America during the Early Triassic to the Middle Triassic period (latest Olenekian to Anisian stages). All species had large "sails" on their backs. Ctenosauriscids are among some of the earliest archosaurs and represent the first global radiation of the group. Genera Phylogeny Ctenosauriscidae was named by Oskar Kuhn in 1964 in paleontology, 1964 to include the genus ''Ctenosauriscus''. It is a stem-based taxon defined by Richard J. Butler, Stephen L. Brusatte, Mike Reich, Sterling J. Nesbitt, Rainer R. Schoch and Jahn J. Hornung in 2011 as "the most inclusive clade containing ''Ctenosauriscus koeneni'' but not ''Poposaurus gracilis'', ''Effigia okeeffeae'', ''Postosuchus kirkpatricki'', ''Crocodylus niloticus'', ''Ornithosuchus longidens'', or ''Aetosaurus ferratus''". The cladograms below follows a 2011 ...
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Tsylmosuchus
''Tsylmosuchus'' is an extinct genus of potentially dubious ctenosauriscid archosauriform reptile known from Western Russia. Fossils referred to ''Tsylmosuchus'' occurred over a wide area in sediments corresponding to the Induan and Olenekian stages of the Early Triassic. Most of these fossils are fragmentary neck vertebrae which were originally reported as sharing similarities with crocodile-line archosaurs (pseudosuchians) such as ''Mandasuchus''. As a result, ''Tsylmosuchus'' was first described as part of the family Rauisuchidae, making it supposedly one of the oldest known archosaurs. However, its fragmentary remains do not show any of the distinguishing features of rauisuchids or even pseudosuchians in general, so ''Tsylmosuchus'' has more recently been interpreted as an indeterminate archosauriform. Although three species of ''Tsylmosuchus'' have been named, they lack diagnostic traits and are probably not distinct from each other. Discovery The genus is named after th ...
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Arizonasaurus
''Arizonasaurus'' was a ctenosauriscid archosaur from the Middle Triassic (243 million years ago). ''Arizonasaurus'' is found in the Middle Triassic Moenkopi Formation of northern Arizona. A fairly complete skeleton was found in 2002 by Sterling Nesbitt. The taxon has a large sailback formed by elongated neural spines of the vertebrae. The type species, ''Arizonasaurus babbitti'', was named by Samuel Paul Welles in 1947. Discovery and naming The type species, ''Arizonasaurus babbitti'', was named by Samuel Paul Welles in 1947 on the basis of a few teeth and a maxilla, labelled as specimen UCMP 36232. A fairly complete skeleton was found in 2002 by Sterling Nesbitt. Description ''Arizonasaurus'' had a sail made of tall neural spines. This sail was similar to those of other basal archosaurs, such as other ctenosauriscids like ''Ctenosauriscus'', ''Lotosaurus'', ''Bromsgroveia'', and ''Hypselorhachis''. ''Arizonasaurus'' is described from two braincase specimens. Some ancestral f ...
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Arizonasaurus BW
''Arizonasaurus'' was a Ctenosauriscidae, ctenosauriscid archosaur from the Middle Triassic (243 million years ago). ''Arizonasaurus'' is found in the Middle Triassic Moenkopi Formation of northern Arizona. A fairly complete skeleton was found in 2002 by Sterling Nesbitt. The taxon has a large sailback formed by elongated neural spines of the vertebrae. The type species, ''Arizonasaurus babbitti'', was named by Samuel Paul Welles in 1947. Discovery and naming The type species, ''Arizonasaurus babbitti'', was named by Samuel Paul Welles in 1947 on the basis of a few teeth and a maxilla, labelled as specimen UCMP 36232. A fairly complete skeleton was found in 2002 by Sterling Nesbitt. Description ''Arizonasaurus'' had a sail made of tall neural spines. This sail was similar to those of other basal archosaurs, such as other ctenosauriscidae, ctenosauriscids like ''Ctenosauriscus'', ''Lotosaurus'', ''Bromsgroveia'', and ''Hypselorhachis''. ''Arizonasaurus'' is described from two br ...
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Early Triassic
The Early Triassic is the first of three epochs of the Triassic Period of the geologic timescale. It spans the time between Ma and Ma (million years ago). Rocks from this epoch are collectively known as the Lower Triassic Series, which is a unit in chronostratigraphy. The Early Triassic is the oldest epoch of the Mesozoic Era. It is preceded by the Lopingian Epoch (late Permian, Paleozoic Era) and followed by the Middle Triassic Epoch. The Early Triassic is divided into the Induan and Olenekian ages. The Induan is subdivided into the Griesbachian and Dienerian subages and the Olenekian is subdivided into the Smithian and Spathian subages. The Lower Triassic series is coeval with the Scythian Stage, which is today not included in the official timescales but can be found in older literature. In Europe, most of the Lower Triassic is composed of Buntsandstein, a lithostratigraphic unit of continental red beds. The Early Triassic and partly also the Middle Triassic span the in ...
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North America
North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and the Caribbean Sea, and to the west and south by the Pacific Ocean. Because it is on the North American Plate, North American Tectonic Plate, Greenland is included as a part of North America geographically. North America covers an area of about , about 16.5% of Earth's land area and about 4.8% of its total surface. North America is the third-largest continent by area, following Asia and Africa, and the list of continents and continental subregions by population, fourth by population after Asia, Africa, and Europe. In 2013, its population was estimated at nearly 579 million people in List of sovereign states and dependent territories in North America, 23 independent states, or about 7.5% of the world's population. In Americas (terminology)#Human ge ...
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Oskar Kuhn
Oskar Kuhn (7 March 1908, Munich – 1990) was a German palaeontologist. Life and career Kuhn was educated in Dinkelsbühl and Bamberg and then studied natural science, specialising in geology and paleontology, at the University of Munich, from which he received his D. Phil. in 1932. He worked in the University of Munich Geological Institute, among other things on the ''Fossilium Catalogus'' (Catalogue of Fossils), and then in 1938 on a stipend from the Deutsche Forschungsgemeinschaft, moved to the University of Halle, where he worked on the Geiseltal fossils. In 1939 he achieved his Habilitation with a thesis on the Halberstadt Keuper fauna, and in 1940 was named Privatdozent in geology and paleontology. Informed by his Catholic religion, Kuhn was an exponent of idealistic morphology: he viewed evolution as operating only within predetermined morphological classes. In 1943 he declared, "The theory of descent has collapsed." After a political conflict with his mentor, Johan ...
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Röt Formation
The Röt Formation or Rötton Formation (German for Röt Shale), or Upper Buntsandstein, is a geologic formation of the Buntsandstein in Germany. It preserves fossils dating back to the Middle Triassic Epoch (Anisian or Aegean or Bithynian in the regional stratigraphy).Röt Formation
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The formation overlies the Plattenstein and Solling Formations and is overlain by the

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Manda Formation
The Manda Formation (also known as the Manda Beds) is a Middle Triassic (Anisian?) or possibly Late Triassic (Carnian?) geologic formation in Tanzania. It preserves fossils of many terrestrial vertebrates from the Triassic, including some of the earliest dinosauromorph archosaurs. The formation is often considered to be Anisian in age according to general tetrapod biochronology hypotheses and correlations to the ''Cynognathus'' Assemblage Zone of South Africa. However, some recent studies cast doubt to this age, suggesting that parts deposits may actually be younger (Carnian) in age. History of study One of the first to study rocks of the Manda Formation was British geologist G. M. Stockley. In 1932, Stockley explored the geology of the Ruhuhu Basin in Tanzania. He called a series of layers dating from the Late Carboniferous to the Middle Triassic the Songea Series and divided it into eight units labelled K1-K8. Stockley was also the first to describe fossils from these rocks, n ...
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Solling Formation
The Solling Formation is a geologic formation in Germany.Solling Formation
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Formerly considered earliest (), but later dating places the formation from the to

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Lipovskaya Formation
The Yarenskian Gorizont (geology), Gorizont ("Yarenskian Horizon") is a major biostratigraphic unit in Russia corresponding to Upper Olenekian-age terrestrial sediments. It lies above the Ustmylian Gorizont of the Vetlugian Supergorizont, and is located below a short unconformity overlain by the Donguz Gorizont. Some paleontologists elevate the Yarenskian to a supergorizont and subdivide it into two smaller units: the older Fedorovskian Gorizont and the younger Gamskian Gorizont. Others prefer to consider the Fedorovskian and Gamskian to be subgorizonts or members of the Yarenskian. Vertebrate index fossils of the Fedorovskian include the lungfish ''Gnathorhiza,'' the Trematosauridae, trematosaurid ''Inflectosaurus,'' and the procolophonid ''Burtensia''. In the Gamskian, these index fossils are replaced by ''Ceratodus'', ''Trematosaurus'', and ''Kapes (genus), Kapes'', respectively. The Yarenskian as a whole is characterized by the capitosaur amphibian ''Parotosuchus'', and has also ...
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Bromsgrove Sandstone Formation
The Sherwood Sandstone Group is a Triassic lithostratigraphic group (a sequence of rock strata) which is widespread in Britain, especially in the English Midlands. The name is derived from Sherwood Forest in Nottinghamshire which is underlain by rocks of this age. It has economic importance as the reservoir of the Morecambe Bay gas field, the second largest gas field in the UK. Geographical extent These rocks are found in northwest England as far north as Carlisle (and extending just into Scotland around Annan and Gretna) and in the Vale of Eden and then extending down the Cumbrian coast into Lancashire and Cheshire. They are mostly obscured by superficial deposits but the highest coastal cliffs in northwest England at St Bees Head are formed in the St Bees Sandstone, the lowermost formation within the group. In the northeast they extend from Hartlepool south through the Vale of Mowbray and the Vale of York, then south through Nottinghamshire into the English Midlands, though ar ...
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