Piloceratidae
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Piloceratidae
The Piloceratidae are a compressed, rapidly expanding, cyrtoconic brevicones with holochoanitic ventral siphuncles and simple endocones. Most likely evolved from ''Clitendoceras'', a narrow, slightly endogastric genus intermediate in form between straight shelled ''Proendoceras'' and the bulkier Piloceratidae. Found in shallow carbonate marine sediments of Demingian through the Cassinian age, (essentially Arenigian, = early Middle -Upper Canadian) . Pilocerids split off from the Proterocameroceratidae very shortly after their inception and are the first family which the proterocamerocerids gave rise to. With the exception of Humeoceras, found in the middle Silurian, pilocerid genera are limited to the Lower Ordovician (Canadian in North America). The general shape of the pilocerid shell precludes an ambush predator lying in wait on the sea floor, or a stealthful hunter drifting through the water. Rather, they probably crawled over the sea floor, head down with the shell off the ...
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Cassinoceras
''Cassinoceras'' ("Cassin Horn") is a genus of nautiloids belonging to the endocerid family Piloceratidae that comes from the late Early Ordovician of eastern North America and adjacent territories.Teichert, Curt, 1964. Endoceratoidea ''in'' Treatise on Invertebrate Paleontology Part K Mollusca 3; Geological Soc. of America and Univ of Kansas Press; pp K171-K172, Piloceratidae. ''Cassinoceras'' is characterized by short, laterally compressed, rapidly expanding shells in which the upper, dorsal, surface has a convex curvature and diverges strongly from the essentially straight lower, ventral surface. The siphuncle which entirely fills the apical portion at the back of the shell is also compressed and vertically expanding, and is filled with simple endocones with a complex system of endosiphuncular blades through which a flattened endosiphuncular tube with widely spaced partitions runs. The shell of ''Cassinoceras'' grew to about 25 to 30 cm long.Moore, Raymond C., Lalicker, C ...
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Proterocameroceratidae
The ''Proterocameroceratidae'' were the first of the Endocerida. They began early in the Ordovician with ''Proendoceras'' or similar genus which had developed endocones, replacing the diaphragms of the ellesmerocerid ancestor. Proterocameroceratids are long, straight or gently curved with a generally narrow siphuncle along the ventral margin. Septal necks are short, never quite reaching the previous septum and may vary in length ontogenically within a species. Connecting rings are thick and layered. Endocones are simple, especially in early forms but may be complex with secondary structures in later forms. The Proterocameroceratidae gave rise to the Piloceratidae early on, and later to the Manchuroceratidae and Chihlioceratidae, from which the Allotrioceratidae are derived, and later yet possibly to the Emmonsoceratidae and Najaceratidae. The Piloceratidae in turn may have given rise to the Endoceratidae although a proterocameroceratid ancestor remains possible. Proterocamerocer ...
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Clitendoceras
''Clitendoceras'' is a genus of cephalopods in the order Endocerida from the Lower Ordovician (m-u Canadian) with an elongate shell with a slight downward, endogastric, curvature and a siphuncle that lies along the ventral margin. Common for endocerids, the chambers are short and the septa close spaced with sutures sloping forward across the back of the shell. Septal necks are short in the young, lengthening in the adult. Endocones are simple, but with the ventral side of the last formed projecting forward. The endosiphotube running down the middle is arched on top and somewhat flat on the lower side. ''Clitendoceras'' appears early in the Domingian stage, Lower Ordovician, probably derived from '' Proendoceras'', and is the likely ancestor of the mostly compressed and breviconic pilocerids as well as of more similar forms such as ''Mcqueenoceras ''Mcqueenoceras'' is an extinct genus of early endocerid, a nautiloid from the Floian epoch of the late early Ordovician period. ...
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Ordovician
The Ordovician ( ) is a geologic period and System (geology), system, the second of six periods of the Paleozoic Era (geology), Era. The Ordovician spans 41.6 million years from the end of the Cambrian Period million years ago (Mya) to the start of the Silurian Period Mya. The Ordovician, named after the Celtic Britons, Welsh tribe of the Ordovices, was defined by Charles Lapworth in 1879 to resolve a dispute between followers of Adam Sedgwick and Roderick Murchison, who were placing the same Rock (geology), rock beds in North Wales in the Cambrian and Silurian systems, respectively. Lapworth recognized that the fossil fauna in the disputed Stratum, strata were different from those of either the Cambrian or the Silurian systems, and placed them in a system of their own. The Ordovician received international approval in 1960 (forty years after Lapworth's death), when it was adopted as an official period of the Paleozoic Era by the International Union of Geological Sciences, Intern ...
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Siphuncle
The siphuncle is a strand of tissue passing longitudinally through the shell of a cephalopod mollusk. Only cephalopods with chambered shells have siphuncles, such as the extinct ammonites and belemnites, and the living nautiluses, cuttlefish, and ''Spirula''. In the case of the cuttlefish, the siphuncle is indistinct and connects all the small chambers of that animal's highly modified shell; in the other cephalopods it is thread-like and passes through small openings in the septa (walls) dividing the camerae (chambers). Some older studies have used the term siphon for the siphuncle, though this naming convention is uncommon in modern studies to prevent confusion with a mollusc organ of the same name. Function The siphuncle is used primarily in emptying water from new chambers as the shell grows. To perform this task, the cephalopod increases the saltiness of the blood in the siphuncle, and the water moves from the more dilute chamber into the blood through osmosis. At the sam ...
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Silurian
The Silurian ( ) is a geologic period and system spanning 24.6 million years from the end of the Ordovician Period, at million years ago ( Mya), to the beginning of the Devonian Period, Mya. The Silurian is the shortest period of the Paleozoic Era. As with other geologic periods, the rock beds that define the period's start and end are well identified, but the exact dates are uncertain by a few million years. The base of the Silurian is set at a series of major Ordovician–Silurian extinction events when up to 60% of marine genera were wiped out. One important event in this period was the initial establishment of terrestrial life in what is known as the Silurian-Devonian Terrestrial Revolution: vascular plants emerged from more primitive land plants, dikaryan fungi started expanding and diversifying along with glomeromycotan fungi, and three groups of arthropods (myriapods, arachnids and hexapods) became fully terrestrialized. A significant evolutionary milestone during ...
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Nautilus
The nautilus (, ) is a pelagic marine mollusc of the cephalopod family Nautilidae. The nautilus is the sole extant family of the superfamily Nautilaceae and of its smaller but near equal suborder, Nautilina. It comprises six living species in two genera, the type of which is the genus ''Nautilus''. Though it more specifically refers to species ''Nautilus pompilius'', the name chambered nautilus is also used for any of the Nautilidae. All are protected under CITES Appendix II. Depending on species, adult shell diameter is between 4 and 10 inches. Nautilidae, both extant and extinct, are characterized by involute or more or less convolute shells that are generally smooth, with compressed or depressed whorl sections, straight to sinuous sutures, and a tubular, generally central siphuncle.Kümmel, B. 1964. Nautiloidae-Nautilida, in the Treatise on Invertebrate Paleontology, Geological Society of America and Univ of Kansas Press, Teichert and Moore eds. Having survived relatively u ...
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