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The ''Proterocameroceratidae'' were the first of the
Endocerida Endocerida is an extinct nautiloid order, a group of cephalopods from the Lower Paleozoic with cone-like deposits in their siphuncle. Endocerida was a diverse group of cephalopods that lived from the Early Ordovician possibly to the Late Silu ...
. They began early in the Ordovician with ''Proendoceras'' or similar genus which had developed endocones, replacing the diaphragms of the ellesmerocerid ancestor. Proterocameroceratids are long, straight or gently curved with a generally narrow siphuncle along the ventral margin. Septal necks are short, never quite reaching the previous septum and may vary in length ontogenically within a species. Connecting rings are thick and layered. Endocones are simple, especially in early forms but may be complex with secondary structures in later forms. The Proterocameroceratidae gave rise to the Piloceratidae early on, and later to the Manchuroceratidae and Chihlioceratidae, from which the Allotrioceratidae are derived, and later yet possibly to the Emmonsoceratidae and Najaceratidae. The Piloceratidae in turn may have given rise to the
Endoceratidae Endoceratidae is a family of large to very large straight shelled nautiloid cephalopods belonging to the order Endocerida that lived during the Middle and Late Ordovician. They include the largest known Paleozoic invertebrates, represented by '' ...
although a proterocameroceratid ancestor remains possible. Proterocameroceratids were probably mobile bottom predators that moved from place to place over the sea floor in search of prey or escape. Their shell was on one hand a carry over from some Cambrian monoplacophoran ancestor, passed on through previous orders, and on the other both a hydrodynamic facilitator and protection. But the shells of nautiloids, such as the proterocameroceratids was also an impediment to true squidlike or fishlike swimming. They could not have had fins for propulsion or steering, only the hyponome for spontaneous jet mobility and their tentacles by which to crawl across the sea floor and to grasp at things. The diagnostic endocones in the early part of the siphuncle balanced the weight of the animal living at the front of the shell. The Proterocameroceratidae ultimately suffered the extinction at the end of the Ordovician, but not before having given rise to a variety of endocerid families. They remain a subject of interesting research.


Genera

*'' Anthoceras'' *'' Campendoceras'' *''
Clitendoceras ''Clitendoceras'' is a genus of cephalopods in the order Endocerida from the Lower Ordovician (m-u Canadian) with an elongate shell with a slight downward, endogastric, curvature and a siphuncle that lies along the ventral margin. Common for end ...
'' *'' Cotteroceras'' *'' Escharendoceras'' *'' Exoclitendoceras'' *'' Lobendoceras'' *'' Lobosiphon'' *'' Manitouoceras'' *'' Mccluskiceras'' *''
Mcqueenoceras ''Mcqueenoceras'' is an extinct genus of early endocerid, a nautiloid from the Floian epoch of the late early Ordovician period. It was similar in overall form to '' Clitendoceras'', from which it may have been derived. ''Mcqueenoceras'', like '' ...
'' *''
Meniscoceras ''Meniscoceras'' is a straight and slender Chazyan endocerid described by Rousseau Flower in 1941 The genus was originally included in the Proterocameroceratidae (Flower, 1955) but later (Flower, 1976) placed with its predecessor, '' Najaceras ...
'' *'' Mysticoceras'' *'' Oderoceras'' *'' Paraendoceras'' *'' Phragmosiphon'' *'' Platysiphon'' *'' Pliendoceras'' *'' Proendoceras'' *'' Proterocameroceras'' *'' Protocyptendoceras'' *'' Retroclitendoceras'' *'' Stenosiphon''


References

*Flower, R. H. 1955, Status of Endoceroid Classification; Jour. Paleon. V 29. n.3 May 1955pp 327–370. * — 1958, Some Chazyan and Mohawkian Endoceratida, Jour Paleon V 32, n 2,May 1858, pp 433–458. * — 1976, Ordovician Cephalopod Faunas and Their Role in Correlation, pp 531–537 in The Ordovician System: Proceedings of a Paleontological Association Symposium; Univ of Wales and Welsh Nat’l Mus Press. *Teichert, Curt 1964, Proterocameroceratidae in the Treatise on Invertebrate Paleontology, VolK pp K166 -k170; Geol Soc of America and University of Kansas Press {{Taxonbar, from=Q7251560 Nautiloids Ordovician cephalopods Ordovician first appearances Late Ordovician extinctions Prehistoric nautiloid families