Pachyrukhinae
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Pachyrukhinae
Hegetotheriidae is an extinct family of notoungulate mammals known from the Oligocene through the Pliocene of South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the southe .... References Typotheres Oligocene mammals Miocene mammals of South America Pliocene mammals Pliocene extinctions Pliocene notoungulates Prehistoric mammal families {{paleo-mammal-stub ...
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Tremacyllus
''Tremacyllus'' is an extinct genus of hegetotheriids. It lived from the Late Miocene to the Late Pleistocene (~7-0.012 Ma) and its fossilized remains were discovered in South America. Description This animal was approximately the size of a hare, and both animals, while unrelated, must have been quite similar in appearance. Its skull had large orbits and strong lower incisors, similar to modern lagomorphs. It was probably a fast animal, with long legs, although proportionally shorter than other similar animals such as ''Pachyrukhos'' or extant lagomorphs. Compared to its relative ''Paedotherium'', ''Tremacyllus'' was slightly smaller and possessed several distinctive characteristics in its dentition: its diastema was longer, the third upper molar was shorter or had the same size than the second molar, and the lower premolars were more overlapping and less molar-like. Furthermore, the symphysis of the mandible was shorter than in ''Paedotherium''. Classification The genus ''Tre ...
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Medistylus
''Medistylus'' is an extinct genus of pachyrukhine hegetotheriid. It lived in Argentina during the Late Oligocene. ''Medistylus'' is known from its upper teeth and isolated skulls, however, its lower dentition is currently unknown. Its fossilized remains were found at the Cabeza Blanca and Las Cascadas localities of the Sarmiento Formation in Chubut Province of Argentina. Description ''Medistylus'' was one of the largest members of the subfamily Pachyrukhinae and had continuously growing teeth, like ''Propachyrucos''. It had a huge pair of front incisors, set obliquely. It lacked the teeth between the first incisor and the second molar, and instead had a toothless space, known as a diastema A diastema (plural diastemata, from Greek διάστημα, space) is a space or gap between two teeth. Many species of mammals have diastemata as a normal feature, most commonly between the incisors and molars. More colloquially, the condition .... It fed on low-lying grass, which caus ...
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Paedotherium
''Paedotherium'' is an extinct, potentially paraphyletic genus of Notoungulate, belonging to the family Hegetotheriidae, composed of small-sized, rodent or lagomorph-like Meridiungulata, South American ungulates. Four species are unambiguously recognized, from the Late Miocene to the Pleistocene of Argentina, and from the late Miocene of Bolivia and Chile. History The first remains associated today with ''Paedotherium'' were first described in 1887 by Florentino Ameghino, and attributed to the related genus ''Pachyruckhos'' due to an important number of similarities. He name the remains ''Pachyrukhos bonaerense''. Its holotype is MACN 1184, remains of the palatal area including the incisors, molars and premolars, to which he include another specimen, MACN 1667, both assumed to come from the Ensenada Formation. In 1888 Hermann Burmeister describe the new genus ''Paedotherium'' insigne. It is, however, rapidly considered a synonym of ''Pachyrukhos'', which take precedence and to ...
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Prosotherium
''Prosotherium'' is an extinct genus of hegetotheriid notoungulate. It lived during the Late Oligocene (between ~29-24 Ma), and its fossilized remains were found in South America. Description This animal was similar to rabbits, in aspect and in size. Its hind legs were particularly long. Crania Its crania was light and thin, notably in the posterior part. The tympanic part of the temporal bone was particularly developed, even more than its relative ''Pachyrukhos'', and it is probable that its auricle was quite large, similar to the ears of a hare. The maxilla and the mandible were high and deep ; the mandible had a subtle coronoid process. The teeth were characterized by incisors pointing inward, molars and premolars covered by a thin layer of dental cementum, generally on the outside part of the upper teeth and on the inside part of the lower teeth. Postcranial skeleton The humerus was large and thin, like the ulna and radius. Metacarpals and phalanges indicates that the ...
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Pachyrukhinae
Hegetotheriidae is an extinct family of notoungulate mammals known from the Oligocene through the Pliocene of South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the southe .... References Typotheres Oligocene mammals Miocene mammals of South America Pliocene mammals Pliocene extinctions Pliocene notoungulates Prehistoric mammal families {{paleo-mammal-stub ...
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Hegetotheriinae
Hegetotheriidae is an extinct family of notoungulate mammals known from the Oligocene through the Pliocene of South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the southe .... References Typotheres Oligocene mammals Miocene mammals of South America Pliocene mammals Pliocene extinctions Pliocene notoungulates Prehistoric mammal families {{paleo-mammal-stub ...
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Propachyrucos
''Propachyrucos'' is an extinct genus of hegetotheriid notoungulate. It lived from the Late Oligocene to the Early Miocene, in what is today South America. Description This animal was the size of a large hare, with a 10 centimeter long skull. Its body was slender, with a very short tail and strong forelegs, although much shorter than the hind legs, equipped with long metatarsals. The skull had a complete dentition, but the second and third incisors, as well as the canines and the first lower premolars, were vestigial. The first upper incisors were very large, and showed a considerable development. The molars had two internal folds separated by a further deep fold, like the Interatheriidae ; dental cementum was present, but to a lesser extent than in ''Interatherium''. The lower teeth were flat on the outside and bilobed on the inside. The tibia and fibula were not fused, and the forearm and lower leg bones were respectively shorter than the humerus and femur, but the hind legs ...
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Hegetotherium
''Hegetotherium'' is an extinct genus of mammals from the Early to Middle Miocene (Colhuehuapian-Colloncuran in the SALMA classification) of South America. Fossils of this genus have been found in the Cerro Bandera, Cerro Boleadoras, Chichinales, Collón Curá, Santa Cruz and Sarmiento Formations of Argentina, the Nazareno Formation of Bolivia, and the Galera and Río Frías Formations of Chile. Taxonomy ''Hegetotherium'' is currently restricted to the type species, ''H. mirable'', of which ''H. convexum'', ''H. anceps'', ''H. minum'' and ''H. andinum'' are synonyms, but also ''H. cerdasensis''. ''"Hegetotherium" arctum'' was formerly assigned to this genus, but is clearly not a member of Hegetotheriidae. ''"Hegetotherium" novum'' was formerly referred to the closely related genus ''Prohegetotherium ''Prohegetotherium'' is an extinct genus of Hegetotheriidae, hegetotheriid Notoungulata, notoungulates from the Oligocene, Late Oligocene to Miocene, Early Miocene (Deseadan ...
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Oligocene
The Oligocene ( ) is a geologic epoch of the Paleogene Period and extends from about 33.9 million to 23 million years before the present ( to ). As with other older geologic periods, the rock beds that define the epoch are well identified but the exact dates of the start and end of the epoch are slightly uncertain. The name Oligocene was coined in 1854 by the German paleontologist Heinrich Ernst Beyrich from his studies of marine beds in Belgium and Germany. The name comes from the Ancient Greek (''olígos'', "few") and (''kainós'', "new"), and refers to the sparsity of extant forms of molluscs. The Oligocene is preceded by the Eocene Epoch and is followed by the Miocene Epoch. The Oligocene is the third and final epoch of the Paleogene Period. The Oligocene is often considered an important time of transition, a link between the archaic world of the tropical Eocene and the more modern ecosystems of the Miocene. Major changes during the Oligocene included a global expansion o ...
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Pliocene Extinctions
The Pliocene ( ; also Pleiocene) is the epoch in the geologic time scale that extends from 5.333 million to 2.58See the 2014 version of the ICS geologic time scale
million years ago. It is the second and most recent epoch of the Period in the . The Pliocene follows the Epoch and is followed by the

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Pliocene Mammals
The Pliocene ( ; also Pleiocene) is the epoch in the geologic time scale that extends from 5.333 million to 2.58See the 2014 version of the ICS geologic time scale
million years ago. It is the second and most recent epoch of the Period in the . The Pliocene follows the Epoch and is followed by the

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Miocene Mammals Of South America
The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recent" because it has 18% fewer modern marine invertebrates than the Pliocene has. The Miocene is preceded by the Oligocene and is followed by the Pliocene. As Earth went from the Oligocene through the Miocene and into the Pliocene, the climate slowly cooled towards a series of ice ages. The Miocene boundaries are not marked by a single distinct global event but consist rather of regionally defined boundaries between the warmer Oligocene and the cooler Pliocene Epoch. During the Early Miocene, the Arabian Peninsula collided with Eurasia, severing the connection between the Mediterranean and Indian Ocean, and allowing a faunal interchange to occur between Eurasia and Africa, including the dispersal of proboscideans into Eurasia. During the late ...
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