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Protohadros
''Protohadros'' (meaning "first hadrosaur") is a genus of herbivorous ornithischian dinosaur from the Late Cretaceous ( Cenomanian stage). Gary Byrd, a part-time palaeontologist, discovered some remains of this euornithopod (ribs and an ungual) during early 1994 at Flower Mound, Denton County, north-central Texas. He informed professional palaeontologist Yuong-Nam Lee of the find, who arranged for the entire preserved fossil to be excavated. It was first reported upon in 1996 by Jason Head of the Dedman College of Humanities and Sciences, Southern Methodist University. The type species ''Protohadros byrdi'' was described and named by Head in 1998. The genus name is derived from Greek πρῶτος, ''protos'', "first", en ἁδρός, ''hadros'', "thick", a reference to the fact that Head considered the species the oldest known hadrosaur. The specific name honours Byrd. The holotype, specimen SMU 74582, of ''Protohadros'', was found in the Woodbine Formation, which date ...
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Hadrosauroidea
Hadrosauroidea is a clade or superfamily of ornithischian dinosaurs that includes the "duck-billed" dinosaurs, or hadrosaurids, and all dinosaurs more closely related to them than to ''Iguanodon''. Their remains have been recovered in Asia, Europe, Africa and the Americas. Many primitive hadrosauroids, such as the Asian ''Probactrosaurus'' and ''Altirhinus'', have traditionally been included in a paraphyly, paraphyletic (unnatural grouping) "Iguanodontidae". With cladistics, cladistic analysis, the traditional Iguanodontidae has been largely disbanded, and probably includes only ''Iguanodon'' and perhaps its closest relatives. Classification The cladogram below follows an analysis by Andrew McDonald, 2012, and shows the position of Hadrosauroidea within Styracosterna. The cladogram below follows an analysis by Wu Wenhao and Pascal Godefroit (2012). Cladogram after Prieto-Marquez and Norell (2010). A phylogenetic analysis performed by Ramírez-Velasco ''et al.'' (2012) foun ...
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Flower Mound, Texas
Flower Mound is an incorporated town located in Denton and Tarrant counties in the U.S. state of Texas. Located northwest of Dallas and northeast of Fort Worth adjacent to Grapevine Lake, the town derives its name from a prominent mound located in the center of town. After settlers used the site for religious camps during the 1840s, the area around Flower Mound was first permanently inhabited in the 1850s; however, residents did not incorporate until 1961. Although an effort to create a planned community failed in the early 1970s, Flower Mound's population increased substantially when Dallas/Fort Worth International Airport opened to the south in 1974. As of the 2020 United States census, the population was 75,956, reflecting a 17% increase over the 64,669 counted in the 2010 census. Of the Texas municipalities that label themselves "towns", Flower Mound has the largest population. Flower Mound was the only town with a population greater than 20,000 in the 2020 census. Flow ...
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Woodbine Formation
The Woodbine Group is a geological formation in east Texas whose strata date back to the Early to Middle Cenomanian age of the Late Cretaceous. It is the producing formation of the giant East Texas Oil Field (also known as the "Black Giant") from which over 5.42 billion barrels of oil have been produced. The Woodbine overlies the Maness Shale, Buda Limestone, or older rocks, and underlies the Eagle Ford Group or Austin Chalk. In outcrop the Woodbine Group has been subdivided into the Lewisville Sandstone, Dexter Sandstone, and/or Pepper Shale formations. Thin-bedded sands of the Woodbine and Eagle Ford are collectively referred to as the "Eaglebine" oil and gas play in the southwestern portion of the East Texas region. Dinosaur and crocodilian remains are among the fossils that have been recovered from the formation.Weishampel, David B; et al. (2004). "Dinosaur distribution (Late Cretaceous, North America)." ''in'' Weishampel, David B.; Dodson, Peter; and Osmólska, Halszka (ed ...
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Late Cretaceous
The Late Cretaceous (100.5–66 Ma) is the younger of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''creta'', the Latin word for the white limestone known as chalk. The chalk of northern France and the white cliffs of south-eastern England date from the Cretaceous Period. Climate During the Late Cretaceous, the climate was warmer than present, although throughout the period a cooling trend is evident. The tropics became restricted to equatorial regions and northern latitudes experienced markedly more seasonal climatic conditions. Geography Due to plate tectonics, the Americas were gradually moving westward, causing the Atlantic Ocean to expand. The Western Interior Seaway divided North America into eastern and western halves; Appalachia and Laramidia. India maintained a northward course towards Asia. In the Southern Hemisphere, Australia and Ant ...
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Type Species
In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen(s). Article 67.1 A similar concept is used for suprageneric groups and called a type genus. In botanical nomenclature, these terms have no formal standing under the code of nomenclature, but are sometimes borrowed from zoological nomenclature. In botany, the type of a genus name is a specimen (or, rarely, an illustration) which is also the type of a species name. The species name that has that type can also be referred to as the type of the genus name. Names of genus and family ranks, the various subdivisions of those ranks, and some higher-rank names based on genus names, have such types.
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Asia
Asia (, ) is one of the world's most notable geographical regions, which is either considered a continent in its own right or a subcontinent of Eurasia, which shares the continental landmass of Afro-Eurasia with Africa. Asia covers an area of , about 30% of Earth's total land area and 8.7% of Earth's total surface area. The continent, which has long been home to the majority of the human population, was the site of many of the first civilizations. Its 4.7 billion people constitute roughly 60% of the world's population. In general terms, Asia is bounded on the east by the Pacific Ocean, on the south by the Indian Ocean, and on the north by the Arctic Ocean. The border of Asia with Europe is a historical and cultural construct, as there is no clear physical and geographical separation between them. It is somewhat arbitrary and has moved since its first conception in classical antiquity. The division of Eurasia into two continents reflects East–West cultural, linguistic, ...
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Iguanodontia
Iguanodontia (the iguanodonts) is a clade of herbivorous dinosaurs that lived from the Middle Jurassic to Late Cretaceous. Some members include ''Camptosaurus'', ''Dryosaurus'', ''Iguanodon'', ''Tenontosaurus'', and the hadrosaurids or "duck-billed dinosaurs". Iguanodontians were one of the first groups of dinosaurs to be found. They are among the best known of the dinosaurs, and were among the most diverse and widespread herbivorous dinosaur groups of the Cretaceous period. Classification Iguanodontia is often listed as an infraorder within a suborder Ornithopoda, though Benton (2004) lists Ornithopoda as an infraorder and does not rank Iguanodontia. Traditionally, iguanodontians were grouped into the superfamily Iguanodontoidea and family Iguanodontidae. However, phylogenetic studies show that the traditional "iguanodontids" are a paraphyletic grade leading up to the hadrosaurs (duck-billed dinosaurs). Groups like Iguanodontoidea are sometimes still used as unranked clades ...
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Hadrosauridae
Hadrosaurids (), or duck-billed dinosaurs, are members of the ornithischian family Hadrosauridae. This group is known as the duck-billed dinosaurs for the flat duck-bill appearance of the bones in their snouts. The ornithopod family, which includes genera such as ''Edmontosaurus'' and ''Parasaurolophus'', was a common group of herbivores during the Late Cretaceous Period. Hadrosaurids are descendants of the Upper Jurassic/Lower Cretaceous iguanodontian dinosaurs and had a similar body layout. Hadrosaurs were among the most dominant herbivores during the Late Cretaceous in Asia and North America, and during the close of the Cretaceous several lineages dispersed into Europe, Africa, South America and Antarctica. Like other ornithischians, hadrosaurids had a predentary bone and a pubic bone which was positioned backwards in the pelvis. Unlike more primitive iguanodonts, the teeth of hadrosaurids are stacked into complex structures known as dental batteries, which acted as effective g ...
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Basal (phylogenetics)
In phylogenetics, basal is the direction of the ''base'' (or root) of a phylogenetic tree#Rooted tree, rooted phylogenetic tree or cladogram. The term may be more strictly applied only to nodes adjacent to the root, or more loosely applied to nodes regarded as being close to the root. Note that extant taxa that lie on branches connecting directly to the root are not more closely related to the root than any other extant taxa. While there must always be two or more equally "basal" clades sprouting from the root of every cladogram, those clades may differ widely in taxonomic rank, Phylogenetic diversity, species diversity, or both. If ''C'' is a basal clade within ''D'' that has the lowest rank of all basal clades within ''D'', ''C'' may be described as ''the'' basal taxon of that rank within ''D''. The concept of a 'key innovation' implies some degree of correlation between evolutionary innovation and cladogenesis, diversification. However, such a correlation does not make a given ca ...
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Quadrate Bone
The quadrate bone is a skull bone in most tetrapods, including amphibians, sauropsids (reptiles, birds), and early synapsids. In most tetrapods, the quadrate bone connects to the quadratojugal and squamosal bones in the skull, and forms upper part of the jaw joint. The lower jaw articulates at the articular bone, located at the rear end of the lower jaw. The quadrate bone forms the lower jaw articulation in all classes except mammals. Evolutionarily, it is derived from the hindmost part of the primitive cartilaginous upper jaw. Function in reptiles In certain extinct reptiles, the variation and stability of the morphology of the quadrate bone has helped paleontologists in the species-level taxonomy and identification of mosasaur squamates and spinosaurine dinosaurs. In some lizards and dinosaurs, the quadrate is articulated at both ends and movable. In snakes, the quadrate bone has become elongated and very mobile, and contributes greatly to their ability to swallow very ...
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Pleurokinesis
Cranial kinesis is the term for significant movement of skull bones relative to each other in addition to movement at the joint between the upper and lower jaw. It is usually taken to mean relative movement between the upper jaw and the braincase. Most vertebrates have some form of kinetic skull. Cranial kinesis, or lack thereof, is usually linked to feeding. Animals which must exert powerful bite forces, such as crocodiles, often have rigid skulls with little or no kinesis, for maximum strength. Animals which swallow large prey whole (snakes), which grip awkwardly shaped food items (parrots eating nuts), or, most often, which feed in the water via suction feeding often have very kinetic skulls, frequently with numerous mobile joints. In the case of mammals, which have akinetic skulls (except for perhaps hares), the lack of kinesis is most likely to be related to the secondary palate, which prevents relative movement. This in turn is a consequence of the need to be able to create ...
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Caririchnium
''Caririchnium'' is an ichnogenus of ornithopod dinosaur footprint, belonging to either derived iguanodonts or basal hadrosauroids. It includes the species ''Caririchnium lotus'' from Lower Cretaceous Jiaguan Formation and ''C''. ''protohadrosaurichnos'' from Upper Cretaceous Woodbine Formation.Yuong-Nam Lee, 1997, "Bird and dinosaur footprints in the Woodbine Formation (Cenomanian), Texas", ''Cretaceous Research'' (1997) 18: 849-864 Specimens are also known from the Lower Cretaceous El Castellar Formation and Camarillas Formations. The trackmacker was probably a large styracosternan related to ''Iguanodon''. See also * List of dinosaur ichnogenera This list of dinosaur ichnogenera is a comprehensive listing of all ichnogenera that have been attributed to dinosaurs, excluding class Aves ( birds, both living and those known only from fossils) and purely vernacular terms. The list includes a ... References Dinosaur trace fossils Ornithopods {{Ornithopod-stub ...
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