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Procolophonomorphs
Procolophonomorpha is an order or clade containing most parareptiles. Many papers have applied various definitions to the name, though most of these definitions have since been considered synonymous with modern parareptile clades such as Ankyramorpha and Procolophonia. The current definition of Procolophonomorpha, as defined by Modesto, Scott, & Reisz (2009), is that of as a stem-based group containing ''Procolophon'' and all taxa more closely related to it than to ''Milleretta''. It constitutes a diverse assemblage that includes a number of lizard-like forms, as well as more diverse types such as the pareiasaurs. Lee 1995, 1996, 1997 argues that turtles evolved from pareiasaurs, but this view is no longer considered likely. Rieppel and deBraga 1996 and deBraga and Rieppel, 1997 argue that turtles evolved from sauropterygians, and there is both molecular and fossil (''Pappochelys'') evidence for the origin of turtles among diapsid reptiles. Classification The following cladogra ...
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Bolosauria
Procolophonomorpha is an order or clade containing most parareptiles. Many papers have applied various definitions to the name, though most of these definitions have since been considered synonymous with modern parareptile clades such as Ankyramorpha and Procolophonia. The current definition of Procolophonomorpha, as defined by Modesto, Scott, & Reisz (2009), is that of as a stem-based group containing ''Procolophon'' and all taxa more closely related to it than to ''Milleretta''. It constitutes a diverse assemblage that includes a number of lizard-like forms, as well as more diverse types such as the pareiasaurs. Lee 1995, 1996, 1997 argues that turtles evolved from pareiasaurs, but this view is no longer considered likely. Rieppel and deBraga 1996 and deBraga and Rieppel, 1997 argue that turtles evolved from sauropterygians, and there is both molecular and fossil (''Pappochelys'') evidence for the origin of turtles among diapsid reptiles. Classification The following cladog ...
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Delorhynchus Cifellii
''Delorhynchus'' is an extinct genus of lanthanosuchoid parareptile known from the late Early Permian (Artinskian age) Garber Formation of Comanche County, Oklahoma. It contains three species: the type species ''D. priscus'' is based on a series of maxillae. The second species to be described, ''D. cifellii'', is known from a larger number of well-preserved skulls and skeletal material. The third species, ''D. multidentatus'', is based on a fragmentary skull with several rows of teeth on its jaw. Discovery The type species, ''D. priscus,'' was first described and named by Richard C. Fox in 1962. The generic name "''Delorhynchus''" is derived from Greek ''rhynchos''/''ρυγχος'', meaning "beak" (a common suffix for extinct reptile genera names). The specific name of the type species ''D. priscus'' is derived from Greek ''πρίσκος'', meaning "ancient" or "venerable" in reference to the fragmentary nature of the known remains. ''D. priscus'' is known from the holotyp ...
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Procolophonia
The Procolophonia are a suborder of herbivorous reptiles that lived from the Middle Permian till the end of the Triassic period. They were originally included as a suborder of the Cotylosauria (later renamed Captorhinida Carroll 1988) but are now considered a clade of Parareptilia. They are closely related to other generally lizard-like Permian reptiles such as the Millerettidae, Bolosauridae, Acleistorhinidae, and Lanthanosuchidae, all of which are included under the Anapsida or "Parareptiles" (as opposed to the Eureptilia). Classification There are two main groups of Procolophonia, the small, lizard-like Procolophonoidea, and the Pareiasauroidea, which include the large, armoured Pareiasauridae. According to the traditional classification of Carroll 1988 as well as recent phylogenetic analyses, smaller groups like Rhipaeosauridae (now a synonym of Nycteroleteridae) and Sclerosauridae are classified with the pareiasaurs and with the procolophonids, respectively. The ...
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Colobomycter Pholeter
''Colobomycter'' is an extinct genus of lanthanosuchoid parareptile known from the Early Permian of Oklahoma. Discovery The type species, ''Colobomycter pholeter'', was first described from fossil remains in 1958, at which time it was believed to represent a synapsid, specifically, a pelycosaur. However, the discovery of new material and reexamination of the holotype led to its reclassification as a member of the Eureptilia. More recent studies indicate that ''Colobomycter'' is properly placed within the amniote clade Parareptilia, as part of the group Lanthanosuchoidea and closely related to the taxon ''Acleistorhinus''. A second species of ''Colobomycter'' was described in 2016, ''Colobomycter vaughni''. No postcranial material is known for ''Colobomycter'', and the skull material referred to the genus has all been recovered from a single locality, the Richards Spur site at the Dolese Brothers Limestone Quarry, 11 kilometers north of Fort Sill, Comanche County, Oklahoma, dati ...
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Feeserpeton Oklahomensis
''Feeserpeton'' is an extinct genus of parareptile from the Early Permian of Richard's Spur, Oklahoma. It is known from a single species, ''Feeserpeton oklahomensis'', which was named in 2012 on the basis of a nearly complete skull. ''Feeserpeton'' is a member of the clade Lanthanosuchoidea and is one of the earliest parareptiles. Description The only known skull of ''Feeserpeton'' is small, but well-fused bones, deep pitting, and worn teeth indicate that the individual was close to maturity when it died. Large eye sockets may indicate that ''Feeserpeton'' was nocturnal. The skull is nearly complete, missing parts of the premaxilla (a bone at the tip of the snout) and the jugal (a bone making up the "cheek" region). A combination of features distinguish ''Feeserpeton'' from other related parareptiles, including a triangular skull, large caniniform teeth in the upper and lower jaws, and postorbital bones behind the eye sockets that are much larger than the nearby squamosal bones ...
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Australothyris Smithi
''Australothyris'' is an extinct genus of basal procolophonomorph parareptile known from the Middle Permian (middle Capitanian stage) of ''Tapinocephalus'' Assemblage Zone, South Africa. The type and only known species is ''Australothyris smithi''. As the most basal member of Procolophonomorpha, ''Australothyris'' helped to contextualize the origin of this major parareptile subgroup. It has been used to support the hypotheses that procolophonomorphs originated in Gondwana and ancestrally possess temporal fenestrae, due to its large and fully enclosed temporal fenestra and South African heritage. It also possessed several unique features, including a high tooth number, long postfrontal, small interpterygoid vacuity, and a specialized interaction between the stapes and quadrate. Discovery Australothyris is known from a single specimen discovered at the Beukesplaas farm by Robert Smith in 1995. The fossil site at the Beukesplaas farm contains a diverse parareptile and synapsid f ...
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Australothyris
''Australothyris'' is an extinct genus of basal procolophonomorph parareptile known from the Middle Permian (middle Capitanian stage) of ''Tapinocephalus'' Assemblage Zone, South Africa. The type and only known species is ''Australothyris smithi''. As the most basal member of Procolophonomorpha, ''Australothyris'' helped to contextualize the origin of this major parareptile subgroup. It has been used to support the hypotheses that procolophonomorphs originated in Gondwana and ancestrally possess temporal fenestrae, due to its large and fully enclosed temporal fenestra and South African heritage. It also possessed several unique features, including a high tooth number, long postfrontal, small interpterygoid vacuity, and a specialized interaction between the stapes and quadrate. Discovery Australothyris is known from a single specimen discovered at the Beukesplaas farm by Robert Smith in 1995. The fossil site at the Beukesplaas farm contains a diverse parareptile and synapsid ...
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Bolosauridae
Bolosauridae is an extinct family of ankyramorph parareptiles known from the latest Carboniferous (Gzhelian) or earliest Permian (Asselian) to the early Guadalupian epoch (latest Roadian stage) of North America, China, Germany, Russia and France. The bolosaurids were unusual for their time period by being bipedal, the oldest known tetrapods to have been so. Their teeth suggest that they were herbivores. The bolosaurids were a rare group and died out without any known descendants. The following cladogram shows the phylogenetic In biology, phylogenetics (; from Greek φυλή/ φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups o ... position of the Bolosauridae, from Johannes Müller, Jin-Ling Li and Robert R. Reisz, 2008. References Procolophonomorphs Permian reptiles Paleozoic reptiles of Asia Paleozoic reptiles of Europe Pa ...
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Ankyramorpha
Ankyramorpha ("anchor forms") is an extinct clade of procolophonomorph parareptiles which lived between the early Cisuralian epoch (middle Sakmarian stage) to the latest Triassic period (latest Rhaetian stage) of Africa, Antarctica, Asia, Australia, Europe, North America and South America.Jalil, N. E., & Janvier, P. (2005). Les pareiasaures (Amniota, Parareptilia) du Permien supérieur du Bassin d’Argana, Maroc. Geodiversitas, 27(1), 35-132. This clade was named in a 1996 parareptile study by Michael deBraga and Robert R. Reisz. They provided the name Ankyramorpha for a newly recognized clade encompassing "the most recent common ancestor of Procolophonia and Lanthanosuchoidea and all its descendants", and this clade name sees continued use among modern parareptile studies. A similar name, Hallucicrania, was provided in an earlier 1995 study by Michael S. Y. Lee, who defined it as the node-based taxon formed by the most recent common ancestor of lanthanosuchids and " ...
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Microleter
''Microleter'' is an extinct genus of basal procolophonomorph parareptiles which lived in Oklahoma during the Early Permian period. The type and only known species is ''Microleter mckinzieorum''. ''Microleter'' is one of several parareptile taxa described from the Richards Spur fissure fills, and can be characterized from its high tooth count, lacrimal/ narial contact, short postfrontal, and slit-like temporal emargination edged by the postorbital, jugal, squamosal, and quadratojugal. Contrary to '' Australothyris'', which had a similar phylogenetic position as a basal procolophonomorph, ''Microleter'' suggests that early parareptile evolution occurred in Laurasia and that multiple lineages developed openings or emarginations in the temporal region. Discovery The only known specimen of ''Microleter'' is a well-preserved skull and lower jaw designated as OMNH 71306, the holotype specimen. It was found at the Dolese Brothers limestone quarry near Richards Spur in Comanch ...
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Lanthanosuchoidea
Lanthanosuchoidea is an extinct superfamily of ankyramorph parareptiles from the middle Pennsylvanian to the middle Guadalupian epoch ( Moscovian - Wordian In the geologic timescale, the Wordian is an age or stage of the Permian. It is the middle of three subdivisions of the Guadalupian Epoch or Series. The Wordian lasted between and million years ago (Ma). It was preceded by the Roadian and f ... stages) of Europe, North America and Asia. It was named by the Russian paleontologist Ivachnenko in 1980, and it contains two families Acleistorhinidae and Lanthanosuchidae. Phylogeny Lanthanosuchoidea is a node-based taxon defined in 1997 as "the most recent common ancestor of ''Lanthanosuchus'', ''Lanthaniscus'', and ''Acleistorhinus''". The cladogram below follows the topology from a 2011 analysis by Ruta ''et al.'' The cladogram below follows the topology from a 2016 analysis by MacDougall ''et al.'' However, the phylogenetic analysis conducted by Cisneros ''et al ...
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Pappochelys
''Pappochelys'' (''παπποχέλυς'' [πάππος (grandfather) + χέλυς (turtle)] meaning "grandfather turtle" in Ancient Greek, Greek) is an extinct genus of diapsid reptile possibly related to turtles. The genus contains only one species, ''Pappochelys rosinae'', from the Middle Triassic of Germany, which was named by paleontologists and Hans-Dieter Sues in 2015 in paleontology, 2015. The discovery of ''Pappochelys'' provides strong support for the placement of turtles within Diapsida, a hypothesis that has long been suggested by molecular data, but never previously by the fossil record. It is morphologically intermediate between the definite stem-turtle ''Odontochelys'' from the Late Triassic of China and ''Eunotosaurus'', a reptile from the Middle Permian of South Africa. Description ''Pappochelys'' had a wide body, small skull, and a long tail that makes up about half of the total body length, which is up to . The skull is pointed with large eye sockets. Several ...
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