Pristerognathus Assemblage Zone
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Pristerognathus Assemblage Zone
The ''Pristerognathus'' Assemblage Zone is a tetrapod assemblage zone or biozone which correlates to the upper Abrahamskraal Formation and lowermost Teekloof Formation, Adelaide Subgroup of the Beaufort Group, a fossiliferous and geologically important geological Group of the Karoo Supergroup in South Africa. The thickest outcrops, reaching not more than , occur just east of Sutherland through to Beaufort West in the south and Victoria West in the north. Exposures are also found west of Colesberg and south of Graaff-Reinet. The ''Pristerognathus'' Assemblage Zone is the third biozone of the Beaufort Group. The name of the biozone refers to '' Pristerognathus polyodon'', a medium-sized carnivorous therocephalian therapsid. It is characterized by the presence of this therocephalian species in association with ''Diictodon feliceps'', a small, fossorial dicynodont whose fossils are especially common in this biozone. History The first fossils to be found in the Beaufort Group rock ...
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Permian
The Permian ( ) is a geologic period and stratigraphic system which spans 47 million years from the end of the Carboniferous Period million years ago (Mya), to the beginning of the Triassic Period 251.9 Mya. It is the last period of the Paleozoic Era; the following Triassic Period belongs to the Mesozoic Era. The concept of the Permian was introduced in 1841 by geologist Sir Roderick Murchison, who named it after the region of Perm in Russia. The Permian witnessed the diversification of the two groups of amniotes, the synapsids and the sauropsids ( reptiles). The world at the time was dominated by the supercontinent Pangaea, which had formed due to the collision of Euramerica and Gondwana during the Carboniferous. Pangaea was surrounded by the superocean Panthalassa. The Carboniferous rainforest collapse left behind vast regions of desert within the continental interior. Amniotes, which could better cope with these drier conditions, rose to dominance in place of their am ...
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Karoo Supergroup
The Karoo Supergroup is the most widespread stratigraphic unit in Africa south of the Kalahari Desert. The supergroup consists of a sequence of units, mostly of nonmarine origin, deposited between the Late Carboniferous and Early Jurassic, a period of about 120 million years. In southern Africa, rocks of the Karoo Supergroup cover almost two thirds of the present land surface, including all of Lesotho, almost the whole of Free State, and large parts of the Eastern Cape, Northern Cape, Mpumalanga and KwaZulu-Natal Provinces of South Africa. Karoo supergroup outcrops are also found in Namibia, Eswatini, Zambia, Zimbabwe and Malawi, as well as on other continents that were part of Gondwana. The basins in which it was deposited formed during the formation and breakup of Pangea.McCarthy, T., Rubridge, B. (2005). ''The Story of Earth and Life.'' pp. 161, 187–241. Struik Publishers, Cape Town The type area of the Karoo Supergroup is the Great Karoo in South Africa, where the most exte ...
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Dicynodont
Dicynodontia is an extinct clade of anomodonts, an extinct type of non-mammalian therapsid. Dicynodonts were herbivorous animals with a pair of tusks, hence their name, which means 'two dog tooth'. Members of the group possessed a horny, typically toothless beak, unique amongst all synapsids. Dicynodonts first appeared in Southern Pangaea during the mid-Permian, ca. 270–260 million years ago, and became globally distributed and the dominant herbivorous animals in the Late Permian, ca. 260–252 Mya. They were devastated by the end-Permian Extinction that wiped out most other therapsids ca. 252 Mya. They rebounded during the Triassic but died out towards the end of that period. They were the most successful and diverse of the non-mammalian therapsids, with over 70 genera known, varying from rat-sized burrowers to elephant-sized browsers. Characteristics The dicynodont skull is highly specialised, light but strong, with the synapsid temporal openings at the rear of the skull ...
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Fossorial
A fossorial () animal is one adapted to digging which lives primarily, but not solely, underground. Some examples are badgers, naked mole-rats, clams, meerkats, and mole salamanders, as well as many beetles, wasps, and bees. Prehistoric evidence The physical adaptation of fossoriality is widely accepted as being widespread among many prehistoric phyla and taxa, such as bacteria and early eukaryotes. Furthermore, fossoriality has evolved independently multiple times, even within a single family. Fossorial animals appeared simultaneously with the colonization of land by arthropods in the late Ordovician period (over 440 million years ago). Other notable early burrowers include ''Eocaecilia'' and possibly ''Dinilysia''. The oldest example of burrowing in synapsids, the lineage which includes modern mammals and their ancestors, is a cynodont, ''Thrinaxodon liorhinus'', found in the Karoo of South Africa, estimated to be 251 million years old. Evidence shows that this ...
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Diictodon
''Diictodon'' is an extinct genus of pylaecephalid dicynodont. These mammal-like synapsids lived during the Late Permian period, approximately 255 million years ago. Fossils have been found in the ''Cistecephalus'' Assemblage Zone of the Madumabisa Mudstone of the Luangwa Basin in Zambia and the ''Tropidostoma'' Assemblage Zone of the Teekloof Formation, ''Tapinocephalus'' Assemblage Zone of the Abrahamskraal Formation, ''Dicynodon'' Assemblage Zone of the Balfour Formation, ''Cistecephalus'' Assemblage Zone of the Middleton or Balfour Formation of South Africa and the Guodikeng Formation of China. Roughly half of all Permian vertebrate specimens found in South Africa are those of ''Diictodon''. This small herbivorous animal was one of the most successful synapsids in the Permian period. Characteristics Appearance ''Diictodon'' had disproportionally large heads that ended in a horny beak. Both males and females had a pair of tusks sticking out from the upper jaw, ...
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Therapsid
Therapsida is a major group of eupelycosaurian synapsids that includes mammals, their ancestors and relatives. Many of the traits today seen as unique to mammals had their origin within early therapsids, including limbs that were oriented more underneath the body, as opposed to the sprawling posture of many reptiles and salamanders. Therapsids evolved from "pelycosaurs", specifically within the Sphenacodontia, more than 279.5 million years ago. They replaced the "pelycosaurs" as the dominant large land animals in the Middle Permian through to the Early Triassic. In the aftermath of the Permian–Triassic extinction event, therapsids declined in relative importance to the rapidly diversifying reptiles during the Middle Triassic. The therapsids include the cynodonts, the group that gave rise to mammals ( Mammaliaformes) in the Late Triassic, around 225 million years ago. Of the non-mammalian therapsids, only cynodonts survived beyond the end of the Triassic, with the only other ...
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Therocephalia
Therocephalia is an extinct suborder of eutheriodont therapsids (mammals and their close relatives) from the Permian and Triassic. The therocephalians ("beast-heads") are named after their large skulls, which, along with the structure of their teeth, suggest that they were carnivores. Like other non-mammalian synapsids, therocephalians were once described as "mammal-like reptiles". Therocephalia is the group most closely related to the cynodonts, which gave rise to the mammals. This relationship takes evidence in a variety of skeletal features. The fossils of therocephalians are numerous in the Karoo of South Africa, but have also been found in Russia, China, Tanzania, Zambia, and Antarctica. Early therocephalian fossils discovered in Middle Permian deposits of South Africa support a Gondwanan origin for the group, which seems to have spread quickly across Earth. Although almost every therocephalian lineage ended during the great Permian–Triassic extinction event, a few represe ...
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Carnivore
A carnivore , or meat-eater (Latin, ''caro'', genitive ''carnis'', meaning meat or "flesh" and ''vorare'' meaning "to devour"), is an animal or plant whose food and energy requirements derive from animal tissues (mainly muscle, fat and other soft tissues) whether through hunting or scavenging. Nomenclature Mammal order The technical term for mammals in the order Carnivora is ''carnivoran'', and they are so-named because most member species in the group have a carnivorous diet, but the similarity of the name of the order and the name of the diet causes confusion. Many but not all carnivorans are meat eaters; a few, such as the large and small cats (felidae) are ''obligate'' carnivores (see below). Other classes of carnivore are highly variable. The Ursids, for example: While the Arctic polar bear eats meat almost exclusively (more than 90% of its diet is meat), almost all other bear species are omnivorous, and one species, the giant panda, is nearly exclusively herbivorous. ...
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Graaff-Reinet
Graaff-Reinet is a town in the Eastern Cape Province of South Africa. It is the oldest town in the province. It is also the sixth-oldest town in South Africa, after Cape Town, Stellenbosch, Simon's Town, Paarl and Swellendam. The town was the centre of a short-lived republic in the late 18th century. The town was a starting point for Great Trek groups led by Gerrit Maritz and Piet Retief and furnished large numbers of the Voortrekkers in 1835–1842. Graaff-Reinet is home to more national monuments than any other town or city in South Africa. It is also known for being a flourishing market for agricultural produce, noted for its mohair industry, and sheep and ostrich farming. History Graaff-Reinet was established by the Dutch East India Company in 1786, after Cape Town in 1652, Stellenbosch in 1679, Paarl in 1687 and Swellendam in 1745. The town is named after then-governor of the Cape Colony, Cornelis Jacob van de Graaff, and his wife. The town was originally established ...
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Colesberg
Colesberg is a town with 17,354 inhabitants in the Northern Cape province of South Africa, located on the main N1 road from Cape Town to Johannesburg. In a sheep-farming area spread over half-a-million hectares, greater Colesberg breeds many of the country's top merinos. It is also renowned for producing high-quality racehorses and many stud farms, including one owned by renowned golfer, Gary Player, are nearby. History Founded in 1830 on an abandoned station of the London Missionary Society, and initially named Toverberg after a nearby hill, it was renamed Colesberg after Sir Galbraith Lowry Cole, then Governor of the Cape Colony. The site of the town lay on one of the well-travelled routes used by traders, hunters and explorers to gain access to the interior. Towerberg or Coleskop is a prominent hill near the town and a landmark easily seen from a distance by travellers. Colesberg saw a large number of battles and skirmishes during the second Anglo-Boer War, and the Col ...
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Victoria West
Victoria West is a town in the central Karoo region of South Africa's Northern Cape province. It is situated on the main N12 route, at an elevation of . It is the seat of the Ubuntu Local Municipality within the Pixley ka Seme District Municipality. History The town was laid out on the bank of the Brakrivier watercourse in 1843, when the Dutch Reformed Church bought the farm Zeekoegat from the estate of J.H. Classens. It was named Victoria in 1844, after Queen Victoria, though amended to Victoria West in 1855 to distinguish it from an Eastern Cape district. In 1859 the town acquired municipal status. Dutch Reformed Church The first services of the Dutch Reformed Church were led by Rev. Colin Fraser of Beaufort West in the “Kerkhuis” on the farm Kapoksfontein. The congregation in Victoria West separated from that in Beaufort West during October 1843 when the first local preacher, Rev. W.E. Krige was ordained. For various practical reasons, they decided not to establish the t ...
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Beaufort West
Beaufort West (Afrikaans: ''Beaufort-Wes''; Xhosa: ''eBhofolo'') is a town in the Western Cape province in South Africa. It is the largest town in the arid Great Karoo region, and is known as the "Capital of the Karoo". It forms part of the Beaufort West Local Municipality, with 34,085 inhabitants in 2011. It is the centre of an agricultural district based mainly on sheep farming, and is a significant town and logistical support hub on the N1 national road. Next door to Beaufort West is the Karoo National Park. Important Permian era fossils have been found in the area to the west of the town. Initially found by David Baird, son of the local magistrate in 1827. The old Town Hall and the Dutch Reformed Church have been declared national monuments. History Beaufort West was the first town to be established in the central Karoo. The town was founded in 1818 and initially named ''Beaufort'' after Henry Somerset, 5th Duke of Beaufort, who was the father of Lord Charles Henry Somerset ...
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