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Poposaur
Poposauroidea is a clade of advanced pseudosuchians ( archosaurs closer to crocodilians than to dinosaurs). It includes poposaurids, shuvosaurids, ctenosauriscids, and other unusual pseudosuchians such as '' Qianosuchus'' and '' Lotosaurus''. However, it excludes most large predatory quadrupedal " rauisuchians" such as rauisuchids and "prestosuchids". Those reptiles are now allied with crocodylomorphs (crocodile ancestors) in a clade known as Loricata, which is the sister taxon to the poposauroids in the clade Paracrocodylomorpha. Although it was first formally defined in 2007, the name "Poposauroidea" has been used for many years. The group has been referred to as Poposauridae by some authors, although this name is often used more narrowly to refer to the family that includes ''Poposaurus'' and its close relatives. Poposauroids went extinct at the end of the Triassic period along with other non-crocodylomorph pseudosuchians. However, they were among the most diverse and long ...
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Paracrocodylomorpha
Paracrocodylomorpha is a clade of pseudosuchian archosaurs. The clade includes the diverse and unusual group Poposauroidea as well as the generally carnivorous and quadrupedal members of Loricata, including modern crocodylians. Paracrocodylomorpha was named by paleontologist J. Michael Parrish in 1993, although the group is now considered to encompass more reptiles than his original definition intended. The most recent definition of Paracrocodylomorpha, as defined by Sterling Nesbitt in 2011, is "the least inclusive clade containing ''Poposaurus'' and ''Crocodylus niloticus'' (the Nile crocodile). Most groups of paracrocodylomorphs became extinct at the end of the Triassic period, with the exception of the crocodylomorphs, from which crocodylians such as crocodiles and alligators evolved in the latter part of the Mesozoic. History and definition Parrish (1993) defined Paracrocodylomorpha as the last common ancestor of "Poposauridae" and Crocodylomorpha and all of its descendants ...
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Rauisuchia
"Rauisuchia" is a paraphyletic group of mostly large and carnivorous Triassic archosaurs. Rauisuchians are a category of archosaurs within a larger group called Pseudosuchia, which encompasses all archosaurs more closely related to crocodilia Crocodilia (or Crocodylia, both ) is an order of mostly large, predatory, semiaquatic reptiles, known as crocodilians. They first appeared 95 million years ago in the Late Cretaceous period ( Cenomanian stage) and are the closest living ...ns than to birds and other dinosaurs. First named in the 1940s, Rauisuchia was a name exclusive to Triassic archosaurs which were generally large (often ), carnivorous, and quadrupedal with a pillar-erect hip posture, though exceptions exist for all of these traits. Rauisuchians, as a traditional Taxonomy, taxonomic group, were considered distinct from other Triassic archosaur groups such as early dinosaurs, Phytosaur, phytosaurs (crocodile-like carnivores), Aetosaur, aetosaurs (armored herbivor ...
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Loricata
Loricata is a clade of archosaur reptiles that includes crocodilians and some of their Triassic relatives, such as ''Postosuchus'' and ''Prestosuchus''. More specifically, Loricata includes Crocodylomorpha (the persistent archosaur subset which crocodilians belong to) and most "rauisuchians", a paraphyletic grade of large terrestrial pseudosuchians which were alive in the Triassic period and ancestral to crocodylomorphs. Loricata is one branch of the group Paracrocodylomorpha; the other branch is the clade Poposauroidea, an unusual collection of strange "rauisuchians" including bipedal, herbivorous, and sail-backed forms. The vast majority of typical "rauisuchians", which were usually quadrupedal predators, occupy basal (early-branching) rungs of Loricata leading up to crocodylomorphs. History and Classification Loricata was initially named in a completely different context by German naturalist Blasius Merrem in his 1820 ''Versuch eines Systems der Amphibien''. Merrem consid ...
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Poposaurus
''Poposaurus'' (" Popo Agie reptile") is an extinct genus of pseudosuchian archosaur from the Late Triassic of the southwestern United States. It belongs to the clade Poposauroidea, an unusual group of Triassic pseudosuchians that includes sail-backed, beaked, and aquatic forms. Fossils have been found in Wyoming, Utah, Arizona, and Texas. Except for the skull, most parts of the skeleton are known. The type species, ''P. gracilis'', was described and named by Maurice Goldsmith Mehl in 1915. A second species, ''P. langstoni'', was originally the type species of the genus ''Lythrosuchus''. Since it was first described, ''Poposaurus'' has been variously classified as a dinosaur, a phytosaur, and a "rauisuchian". Like theropod dinosaurs, ''Poposaurus'' was an obligate biped, meaning that it walked on two legs rather than four. However, as a pseudosuchian, it is more closely related to living crocodilians than to dinosaurs. ''Poposaurus'' is thought to have evolved this form of locomo ...
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Xilousuchus
''Xilousuchus'' is an extinct genus of poposauroid from lower Triassic (Olenekian stage) deposits of Fugu County of northeastern Shanxi Province, China. It is known from the holotype, IVPP V 6026, a single well-preserved partial skeleton including the skull. It was found from the Heshanggou Formation of the Ordos Basin, Hazhen commune. It was first named by Xiao-Chun Wu in 1981 and the type species is ''Xilousuchus sapingensis''. Wu (1981) referred ''Xilousuchus'' to the Proterosuchia. Gower and Sennikov (1996) found it to be an erythrosuchian based strictly on the braincase. A more detailed re-description of the genus was provided by Nesbitt ''et al.'' (2010) and found poposauroid affinities. In his massive revision of archosaurs which included a large cladistic analysis, Sterling J. Nesbitt (2011) found ''Xilousuchus'' to be a poposauroid which is most closely related to ''Arizonasaurus''. ''Xilousuchus'' is the oldest archosaur to date, although ''Ctenosauriscus'' ...
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Prestosuchid
Prestosuchidae (in its widest usage) is a polyphyletic grouping of carnivorous archosaurs that lived during the Triassic. They were large active terrestrial apex predators, ranging from around in length. They succeeded the Erythrosuchidae as the largest archosaurs of their time. While resembling erythrosuchids in size and some features of the skull and skeleton, they were more advanced in their erect posture and crocodile-like ankle, indicating more efficient gait. "Prestosuchids" flourished throughout the whole of the middle, and the early part of the late Triassic, and fossils are so far known from Europe, India, Africa (Tanzania), Argentina, and Paleorrota in Brazil. However, for a long time experts disagree regarding the phylogenetic relationships of the group, what genera should be included, and whether indeed the "Prestosuchidae" constitute a distinct family. In 2011, Prestosuchidae in its broadest definition was determined to be a poorly-diagnosed and obsolete polyphylet ...
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Rauisuchid
Rauisuchidae is a group of large (up to or more) predatory Triassic archosaurs. There is some disagreement over which genera should be included in Rauisuchidae and which should be in the related Prestosuchidae and Poposauridae, and indeed whether these should even be thought of as separate valid Family (biology), families. Rauisuchids occurred throughout much of the Triassic, and may have first occurred in the Early Triassic if some archosaurian taxa such as ''Scythosuchus'' and ''Tsylmosuchus'' are considered to be within the family. An early cladistic analysis of crocodylotarsan (pseudosuchian) archosaurs included ''Lotosaurus'', ''Fasolasuchus'', ''Rauisuchus'', and "the Kupferzell rauisuchid" (later called ''Batrachotomus'') within Rauisuchidae.Parrish JM. 1993. Phylogeny of the Crocodylotarsi, with reference to archosaurian and crurotarsan monophyly. ''Journal of Vertebrate Paleontology'' 13: 287-308. However, a later study found that ''Batrachotomus'' was a more Basal (phyl ...
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Poposaurid
Poposauridae is a family of large carnivorous archosaurs which lived alongside dinosaurs during the Late Triassic. They were around long. Poposaurids are known from fossil remains from North and South America. While originally believed to be theropod dinosaurs (they mirrored the theropods in a number of respects, such as features of the skull and bipedal locomotion), cladistic analysis has shown them to be more closely related to crocodiles. An early cladistic analysis of crocodylotarsan archosaurs included ''Poposaurus'', ''Postosuchus'', ''Teratosaurus'', and ''Bromsgroveia'' within Poposauridae.Parrish JM. 1993. Phylogeny of the Crocodylotarsi, with reference to archosaurian and crurotarsan monophyly. ''Journal of Vertebrate Paleontology'' 13: 287-308. However, later studies found ''Teratosaurus'' to be a rauisuchid.Long RA, Murry PA. 1995. Late Triassic (Carnian and Norian) tetrapods from the Southwestern United States. ''New Mexico Museum of Natural History & Science Bul ...
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Lotosaurus
''Lotosaurus'' is an extinct genus of sail-backed poposauroid Poposauroidea is a clade of advanced pseudosuchians (archosaurs closer to crocodilians than to dinosaurs). It includes poposaurids, shuvosaurids, ctenosauriscids, and other unusual pseudosuchians such as ''Qianosuchus'' and ''Lotosaurus''. Howeve ... known from Hunan Province of central China. Discovery ''Lotosaurus'' is known from the holotype IVPP V 4881 (or possibly V 4880), an articulated and well-preserved skeleton. Other referred specimens include IVPP V 48013 (a skull) as well as many articulated and disarticulated skeletal remains concentrated in a bonebed which is almost completely composed of ''Lotosaurus'' bones. All known specimens of this genus were collected from this bonebed, known as the ''Lotosaurus'' site, which belongs to the Batung Formation (or alternatively Xinlingzhen Formation of the Badong Group). Further excavations in 2018 revealed many more specimens, as well as ...
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Poposauridae
Poposauridae is a family of large carnivorous archosaurs which lived alongside dinosaurs during the Late Triassic. They were around long. Poposaurids are known from fossil remains from North and South America. While originally believed to be theropod dinosaurs (they mirrored the theropods in a number of respects, such as features of the skull and bipedal locomotion), cladistic analysis has shown them to be more closely related to crocodiles. An early cladistic analysis of crocodylotarsan archosaurs included ''Poposaurus'', ''Postosuchus'', ''Teratosaurus'', and ''Bromsgroveia'' within Poposauridae.Parrish JM. 1993. Phylogeny of the Crocodylotarsi, with reference to archosaurian and crurotarsan monophyly. ''Journal of Vertebrate Paleontology'' 13: 287-308. However, later studies found ''Teratosaurus'' to be a rauisuchid.Long RA, Murry PA. 1995. Late Triassic (Carnian and Norian) tetrapods from the Southwestern United States. ''New Mexico Museum of Natural History & Science Bul ...
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Effigia
''Effigia'' was an extinct genus of shuvosaurid known from the Late Triassic of New Mexico, south-western USA. With a bipedal stance, long neck, and a toothless beaked skull, ''Effigia'' and other shuvosaurids bore a resemblance to the ornithomimid dinosaurs of the Cretaceous Period. However, shuvosaurids were not dinosaurs, but were instead a specialized family of poposauroid pseudosuchians, meaning that their closest living relatives are crocodilians. Discovery The 2-meter (6 ft 7 in) holotype fossil was collected by Edwin H. Colbert in 1947. At this time Colbert led an excavation to collect blocks of rock from the Whitaker Quarry of Ghost Ranch, near Abiquiu, New Mexico. Colbert's expedition intended to recover abundant fossils of the basal theropod dinosaur ''Coelophysis'', and he believed that no other large vertebrates were present in the quarry. As a result, his crew did not even open the plaster jackets of most of the blocks that were shipped to the American Museum of ...
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Bipedalism
Bipedalism is a form of terrestrial locomotion where an organism moves by means of its two rear limbs or legs. An animal or machine that usually moves in a bipedal manner is known as a biped , meaning 'two feet' (from Latin ''bis'' 'double' and ''pes'' 'foot'). Types of bipedal movement include walking, running, and hopping. Several groups of modern species are habitual bipeds whose normal method of locomotion is two-legged. In the Triassic period some groups of archosaurs (a group that includes crocodiles and dinosaurs) developed bipedalism; among the dinosaurs, all the early forms and many later groups were habitual or exclusive bipeds; the birds are members of a clade of exclusively bipedal dinosaurs, the theropods. Within mammals, habitual bipedalism has evolved multiple times, with the macropods, kangaroo rats and mice, springhare, hopping mice, pangolins and hominin apes ( australopithecines, including humans) as well as various other extinct groups evolving the tra ...
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