|
Placoderms
Placodermi (from Greek πλάξ 'plate' and δέρμα 'skin', literally 'Plate (animal anatomy), plate-skinned') is a Class (biology), class of armoured prehistoric fish, known from fossils, which lived from the Silurian to the end of the Devonian period. Their head and thorax were covered by articulated armoured plates and the rest of the body was scale (zoology), scaled or naked, depending on the species. Placoderms were among the first jawed fish; their Fish jaw, jaws likely evolved from the first of their gill arches. Placoderms are thought to be paraphyly, paraphyletic, consisting of several distinct Outgroup (cladistics), outgroups or sister taxon, sister taxa to all living jawed vertebrates, which originated among their ranks. In contrast, one 2016 analysis concluded that placodermi are likely monophyletic, though these analyses have been further dismissed with more transitional taxa between placoderms and modern gnathosthomes, solidifying their paraphyletic status. Plac ... [...More Info...] [...Related Items...] OR: [Wikipedia] [Google] [Baidu] |
Prehistoric Fish
The evolution of fish began about 530 million years ago during the Cambrian explosion. It was during this time that the early chordates developed the skull and the vertebral column, leading to the first craniates and vertebrates. The first fish lineages belong to the Agnatha, or jawless fish. Early examples include ''Haikouichthys''. During the late Cambrian, eel-like jawless fish called the conodonts, and small mostly armoured fish known as ostracoderms, first appeared. Most jawless fish are now extinct; but the extant lampreys may approximate ancient pre-jawed fish. Lampreys belong to the Cyclostomata, which includes the extant hagfish, and this group may have split early on from other agnathans. The earliest Gnathostomata, jawed vertebrates probably developed during the late Ordovician period. They are first represented in the fossil record from the Silurian by two groups of fish: the armoured fish known as Placodermi, placoderms, which evolved from the ostracoderms; and the ... [...More Info...] [...Related Items...] OR: [Wikipedia] [Google] [Baidu] |
Bianchengichthys
''Bianchengichthys'' is an extinct genus of maxillate placoderm fish from the late Silurian Period. Its fossils have been recovered from Yunnan Province, China, and it is represented by only one species: ''Bianchengichthys micros''. Description ''Bianchengichthys'' is a small, somewhat dorsoventrally compressed placoderm fish. The mandible (made from dermal bone) of this genus differs from ''Entelognathus'' and ''Qilinyu''—two other maxillate placoderms from late Silurian China−in that the oral lamina is broad and carries a row of tooth-like denticles, though the marginal flange is toothless. The pectoral fin, preceded by two small spines on its thoracic shield, is lobate in shape and situated along by a 'fringe' of scales similar to those of ''Lepidotrichia'' in bony fishes. Similarly to other maxillate placoderms, its eyes are anteriorly orientated and very close to its mouth. Evolutionary significance ''Bianchengichthys mandible bears physical resemblance to both its r ... [...More Info...] [...Related Items...] OR: [Wikipedia] [Google] [Baidu] |
Ptyctodontida
The ptyctodontids ("folded-teeth") are placoderms of the order Ptyctodontida, containing the family Ptyctodontidae. With their big heads, big eyes, reduced armor and long bodies, the ptyctodontids bore a superficial resemblance to modern day chimaeras (Holocephali). Their armor was reduced to a pattern of small plates around the head and neck. Like the extinct and related acanthothoracids, and the living and unrelated holocephalians, most of the ptyctodontids are thought to have lived near the sea bottom and preyed on shellfish. On account of their radically reduced armor, some paleontologists have suggested that the Ptyctodontida were not actually placoderms, but actual holocephalians, some primitive group of elasmobranch fish, or even were the ancestors of the holocephalians, including the chimaeras. Thorough anatomical examinations of whole fossil specimens reveal that the profound similarities between these two groups are actually very superficial. The major differences ... [...More Info...] [...Related Items...] OR: [Wikipedia] [Google] [Baidu] |
Bothriolepis
''Bothriolepis'' (from el, βόθρος , 'trench' and el, λεπίς 'scale') was a widespread, abundant and diverse genus of antiarch placoderms that lived during the Middle Devonian, Middle to Late Devonian period of the Paleozoic Era. Historically, ''Bothriolepis'' resided in an array of paleo-environments spread across every paleocontinent, including near shore marine and freshwater settings. Most species of ''Bothriolepis'' were characterized as relatively small, benthic, freshwater detritivores (organisms that obtain nutrients by consuming decomposing plant/animal material), averaging around in length. However, the largest species, ''B. rex'', had an estimated bodylength of . Although expansive with over 60 species found worldwide, comparatively ''Bothriolepis'' is not unusually more diverse than most modern bottom dwelling species around today. Classification ''Bothriolepis'' is a genus placed within the placoderm order Antiarchi. The earliest antiarch placoderms fir ... [...More Info...] [...Related Items...] OR: [Wikipedia] [Google] [Baidu] |
Acanthothoraci
Acanthothoraci (''spine chests'') is an extinct group of chimaera-like placoderms who were closely related to the rhenanid placoderms. Superficially, the acanthoracids resembled scaly chimaeras, or (relatively) heavily armored ptyctodonts. They were distinguished from chimaeras by the presence of large scales and plates, a pair of large spines that emanate from their chests (thus, the order's name), tooth-like beak plates, and the typical bone-enhanced placoderm eyeball. They were distinguished from other placoderms due to differences in the anatomy of their skulls, and due to patterns on the skull plates and thoracic plates that are unique to this order. Fossil record Fossils of the Acanthothoracids are found in various deposits from the Lower Devonian throughout the world. Fossils of the Palaeacanthaspids are found in Eurasia and Canada, while the Weejasperaspids have only been found in the Taemas Wee Jasper reef, in Southeastern Australia. Ecology From what can be inferr ... [...More Info...] [...Related Items...] OR: [Wikipedia] [Google] [Baidu] |
Arthrodira
Arthrodira (Greek for "jointed neck") is an order of extinct armored, jawed fishes of the class Placodermi that flourished in the Devonian period before their sudden extinction, surviving for about 50 million years and penetrating most marine ecological niches. Arthrodires were the largest and most diverse of all groups of Placoderms. Description Arthrodire placoderms are notable for the movable joint between armor surrounding their heads and bodies. Like all placoderms, they lacked distinct teeth; instead, they used the sharpened edges of a bony plate on their jawbone as a biting surface. The eye sockets are protected by a bony ring, a feature shared by birds and some ichthyosaurs. Early arthrodires, such as the genus ''Arctolepis'', were well-armoured fishes with flattened bodies. The largest member of this group, ''Dunkleosteus'', was a true superpredator of the latest Devonian period, reaching as much as 6 m in length. In contrast, the long-nosed ''Rolfosteus'' measured just ... [...More Info...] [...Related Items...] OR: [Wikipedia] [Google] [Baidu] |
Phyllolepida
Phyllolepida ("leaf scales") is an extinct taxon of flattened placoderms found throughout the world, with fossils being found in Devonian strata. Like other flattened placoderms, the phyllolepids were bottom-dwelling predators that ambushed prey. Unlike other flattened placoderms, the phyllolepids were inhabitants of freshwater environments. Unlike the Rhenanida, the armor of the phyllolepids were made of whole plates, rather than numerous tubercles and scales, and unlike the Petalichthyida, the components of the comparatively wide mouth are known. The phyllolepids are considered to have been blind, as the orbits for the eyes are extremely small, so much so as to suggest that the eyes were vestigial, and that they were placed on the sides of the head, as opposed to visually-oriented bottom-dwelling predators, like, say stargazers or flatfish, which have the eyes placed high on top of the head. Despite having a relatively clear idea of the phyllolepids' lifestyle and anatomy, most ... [...More Info...] [...Related Items...] OR: [Wikipedia] [Google] [Baidu] |
Stensioella
''Stensioella heintzi'' is an enigmatic placoderm of arcane affinity. It is only known from the Lower Devonian Hunsrück slate of Germany. Anatomy ''Stensioella heintzi'' has an elongated body, a whip-like tail, and long, wing-like pectoral fins. In life, the animal would have looked vaguely like an elongated ratfish. Like the sympatric ''Gemuendina'', ''S. heintzi'' had armor made up of a complex mosaic of small, scale-like tubercles. Taxonomy ''Stensioella'' is tentatively placed within Placodermi as being among the most basal of all placoderms, as from what can be discerned from the only whole specimen found, the shoulder joints of its armor appear to be very similar to other placoderms. Despite this detail, coupled with superficial similarities in skull plates, and gross, superficial similarities between its tubercles, and the tubercles of the rhenanids, some paleontologists believe that there are very few concrete reasons for ''S. heintzis placement in Placodermi. The ... [...More Info...] [...Related Items...] OR: [Wikipedia] [Google] [Baidu] |
Gnathostomata
Gnathostomata (; from Greek: (') "jaw" + (') "mouth") are the jawed vertebrates. Gnathostome diversity comprises roughly 60,000 species, which accounts for 99% of all living vertebrates, including humans. In addition to opposing jaws, living gnathostomes have true teeth (a characteristic which has subsequently been lost in some), paired appendages (pectoral and pelvic fins, arms, legs, wings, etc.), the elastomeric protein of elastin, and a horizontal semicircular canal of the inner ear, along with physiological and cellular anatomical characters such as the myelin sheaths of neurons, and an adaptive immune system that has the discrete lymphoid organs of spleen and thymus, and uses V(D)J recombination to create antigen recognition sites, rather than using genetic recombination in the variable lymphocyte receptor gene. It is now assumed that Gnathostomata evolved from ancestors that already possessed a pair of both pectoral and pelvic fins. Until recently these ancestors, know ... [...More Info...] [...Related Items...] OR: [Wikipedia] [Google] [Baidu] |
Devonian
The Devonian ( ) is a geologic period and system of the Paleozoic era, spanning 60.3 million years from the end of the Silurian, million years ago (Mya), to the beginning of the Carboniferous, Mya. It is named after Devon, England, where rocks from this period were first studied. The first significant adaptive radiation of life on dry land occurred during the Devonian. Free-sporing vascular plants began to spread across dry land, forming extensive forests which covered the continents. By the middle of the Devonian, several groups of plants had evolved leaves and true roots, and by the end of the period the first seed-bearing plants appeared. The arthropod groups of myriapods, arachnids and hexapods also became well-established early in this period, after starting their expansion to land at least from the Ordovician period. Fish reached substantial diversity during this time, leading the Devonian to often be dubbed the Age of Fishes. The placoderms began dominating ... [...More Info...] [...Related Items...] OR: [Wikipedia] [Google] [Baidu] |
Rhenanida
Rhenanida ("Rhine (fish)") is an Order (biology), order of scaly placoderms. Unlike most other placoderms, the rhenanids' armor was made up of a mosaic of unfused scales and tubercles. The patterns and components of this "mosaic" correspond to the plates of armor in other, more advanced placoderms, suggesting that the ancestral placoderm had armor made of unfused components, as well. All rhenanids were flattened, Batoidea, ray-like, bottom-dwelling predators that lived in marine environments. Evolution The rhenanids were once presumed to be the most primitive, or at least the closest to the ancestral placoderm, as their armor was made up of a mosaic of tubercles, as opposed to the solidified plates of "advanced" placoderms, such as antiarchs and arthrodires. Through comparing the skull anatomies of ''Jagorina pandora'' with those of antiarchs, the rhenanids are considered to be the sister group of the antiarchs (together with their respective Acanthothoracid relatives). Presen ... [...More Info...] [...Related Items...] OR: [Wikipedia] [Google] [Baidu] |
Brindabellaspis
''Brindabellaspis stensioi'' ("Erik Stensiö's Brindabella Ranges Shield") is a placoderm with a flat, platypus-like snout from the Early Devonian of the Taemas-Wee Jasper reef in Australia. When it was first discovered in 1980, it was originally regarded as a Weejasperaspidae, Weejasperaspid acanthothoracid due to anatomical similarities with the other species found at the reef. According to Philippe Janvier, anatomical similarities of ''B. stensiois brain and braincase with those of jawless fish, such as the Osteostraci and the Galeaspida, strongly suggest that ''B. stensioi'', and also Antiarchi, the antiarchs, are basal placoderms closest to the ancestral placoderm. References *Gavin Young, Young, Gavin C. 1980, A new Early Devonian placoderm from New South Wales, Australia, with a discussion of placoderm phylogeny: ''Palaeontographica'' 167A pp. 10–76. 2 pl., 27 fig. * Philippe Janvier, Janvier, Philippe. Early Vertebrates Oxford, New York: Oxford University Press, 1 ... [...More Info...] [...Related Items...] OR: [Wikipedia] [Google] [Baidu] |
