Pholidorhynchodon Malzanni
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Pholidorhynchodon Malzanni
''Pholidorhynchodon'' is an extinct genus of ray-finned fish that lived in the Triassic. Its fossils have been found in Italy, in the Zorzino Limestone Formation in Cene. Significance ''Pholidorhynchodon'' belongs to a group of fish that are right at the very base of the teleosts. Teleost fish include almost all living fish species, and have their origins in the Triassic. This makes ''Pholidorhynchodon'' important because it provides information about the early evolution and development of this important group.Arratia, G., 2017New Triassic teleosts (Actinopterygii, Teleosteomorpha) from northern Italy and their phylogenetic relationships among the most basal teleosts ''Journal of Vertebrate Paleontology'', 37(2), p.e1312690 See also * Prehistoric fish * List of prehistoric bony fish A ''list'' is any set of items in a row. List or lists may also refer to: People * List (surname) Organizations * List College Albert A. List College of Jewish Studies, known simply a ...
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Oligocene
The Oligocene ( ) is a geologic epoch of the Paleogene Period and extends from about 33.9 million to 23 million years before the present ( to ). As with other older geologic periods, the rock beds that define the epoch are well identified but the exact dates of the start and end of the epoch are slightly uncertain. The name Oligocene was coined in 1854 by the German paleontologist Heinrich Ernst Beyrich from his studies of marine beds in Belgium and Germany. The name comes from the Ancient Greek (''olígos'', "few") and (''kainós'', "new"), and refers to the sparsity of extant forms of molluscs. The Oligocene is preceded by the Eocene Epoch and is followed by the Miocene Epoch. The Oligocene is the third and final epoch of the Paleogene Period. The Oligocene is often considered an important time of transition, a link between the archaic world of the tropical Eocene and the more modern ecosystems of the Miocene. Major changes during the Oligocene included a global expansion o ...
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Animalia
Animals are multicellular, eukaryotic organisms in the biological kingdom Animalia. With few exceptions, animals consume organic material, breathe oxygen, are able to move, can reproduce sexually, and go through an ontogenetic stage in which their body consists of a hollow sphere of cells, the blastula, during embryonic development. Over 1.5 million living animal species have been described—of which around 1 million are insects—but it has been estimated there are over 7 million animal species in total. Animals range in length from to . They have complex interactions with each other and their environments, forming intricate food webs. The scientific study of animals is known as zoology. Most living animal species are in Bilateria, a clade whose members have a bilaterally symmetric body plan. The Bilateria include the protostomes, containing animals such as nematodes, arthropods, flatworms, annelids and molluscs, and the deuterostomes, containing the echinode ...
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Chordata
A chordate () is an animal of the phylum Chordata (). All chordates possess, at some point during their larval or adult stages, five synapomorphies, or primary physical characteristics, that distinguish them from all the other taxa. These five synapomorphies include a notochord, dorsal hollow nerve cord, endostyle or thyroid, pharyngeal slits, and a post-anal tail. The name “chordate” comes from the first of these synapomorphies, the notochord, which plays a significant role in chordate structure and movement. Chordates are also Bilateral symmetry, bilaterally symmetric, have a coelom, possess a circulatory system, and exhibit Metameric, metameric segmentation. In addition to the morphological characteristics used to define chordates, analysis of genome sequences has identified two conserved signature indels (CSIs) in their proteins: cyclophilin-like protein and mitochondrial inner membrane protease ATP23, which are exclusively shared by all vertebrates, tunicates and cep ...
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Actinopterygii
Actinopterygii (; ), members of which are known as ray-finned fishes, is a class of bony fish. They comprise over 50% of living vertebrate species. The ray-finned fishes are so called because their fins are webs of skin supported by bony or horny spines (rays), as opposed to the fleshy, lobed fins that characterize the class Sarcopterygii (lobe-finned fish). These actinopterygian fin rays attach directly to the proximal or basal skeletal elements, the radials, which represent the link or connection between these fins and the internal skeleton (e.g., pelvic and pectoral girdles). By species count, actinopterygians dominate the vertebrates, and they constitute nearly 99% of the over 30,000 species of fish. They are ubiquitous throughout freshwater and marine environments from the deep sea to the highest mountain streams. Extant species can range in size from ''Paedocypris'', at , to the massive ocean sunfish, at , and the long-bodied oarfish, at . The vast majority of Actinopt ...
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Teleostei
Teleostei (; Ancient Greek, Greek ''teleios'' "complete" + ''osteon'' "bone"), members of which are known as teleosts ), is, by far, the largest class (biology), infraclass in the class Actinopterygii, the ray-finned fishes, containing 96% of all neontology, extant species of fish. Teleosts are arranged into about 40 order (biology), orders and 448 family (biology), families. Over 26,000 species have been described. Teleosts range from giant oarfish measuring or more, and ocean sunfish weighing over , to the minute male anglerfish ''Photocorynus spiniceps'', just long. Including not only torpedo-shaped fish built for speed, teleosts can be flattened vertically or horizontally, be elongated cylinders or take specialised shapes as in anglerfish and seahorses. The difference between teleosts and other bony fish lies mainly in their jaw bones; teleosts have a movable premaxilla and corresponding modifications in the jaw musculature which make it possible for them to cranial kinesi ...
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Pholidophoridae
Pholidophoridae is an extinct family of primitive stem-teleost fish that lived during the Triassic period. Most of the genera are from the Late Triassic (Carnian and Norian ages) of Europe, but one ('' Malingichthys'') is known from the Middle Triassic (Ladinian) of China. The pholidophorids were historically united with several other stem-teleost lineages into the order Pholidophoriformes; however, Pholidophoriformes in its traditional sense is now thought to be paraphyletic with respect to crown group teleosts. Phylogeny Below is a cladogram A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to d ... simplified after Bean (2021): References Prehistoric ray-finned fish families Ladinian first appearances Norian extinctions Taxa named by Arthur Smith Woodward {{Tr ...
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Triassic
The Triassic ( ) is a geologic period and system which spans 50.6 million years from the end of the Permian Period 251.902 million years ago ( Mya), to the beginning of the Jurassic Period 201.36 Mya. The Triassic is the first and shortest period of the Mesozoic Era. Both the start and end of the period are marked by major extinction events. The Triassic Period is subdivided into three epochs: Early Triassic, Middle Triassic and Late Triassic. The Triassic began in the wake of the Permian–Triassic extinction event, which left the Earth's biosphere impoverished; it was well into the middle of the Triassic before life recovered its former diversity. Three categories of organisms can be distinguished in the Triassic record: survivors from the extinction event, new groups that flourished briefly, and other new groups that went on to dominate the Mesozoic Era. Reptiles, especially archosaurs, were the chief terrestrial vertebrates during this time. A specialized subgroup of archo ...
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Zorzino Limestone Formation
The Calcare di Zorzino, Italian for Zorzino Limestone is a Late Triassic (Norian) geological formation in Italy (Cene and ).Zorzino Limestone
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and have been recovered in this formation.


Vertebrate paleofauna


Other reptiles

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Cene, Lombardy
Cene (Bergamasque: ) is a ''comune'' (municipality) in the Province of Bergamo in the Italian region of Lombardy, located about northeast of Milan and about northeast of Bergamo. Cene borders the following municipalities: Albino, Bianzano, Casnigo, Cazzano Sant'Andrea, Fiorano al Serio, Gaverina Terme, Gazzaniga, Leffe Leffe () is a premium beer brand owned by InBev Belgium, the European operating arm of the global Anheuser–Busch InBev brewery giant. There are several beers in the range, and they are marketed as abbey beers. They are brewed in large quan .... References External links Official website
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Teleosts
Teleostei (; Greek ''teleios'' "complete" + ''osteon'' "bone"), members of which are known as teleosts ), is, by far, the largest infraclass in the class Actinopterygii, the ray-finned fishes, containing 96% of all extant species of fish. Teleosts are arranged into about 40 orders and 448 families. Over 26,000 species have been described. Teleosts range from giant oarfish measuring or more, and ocean sunfish weighing over , to the minute male anglerfish ''Photocorynus spiniceps'', just long. Including not only torpedo-shaped fish built for speed, teleosts can be flattened vertically or horizontally, be elongated cylinders or take specialised shapes as in anglerfish and seahorses. The difference between teleosts and other bony fish lies mainly in their jaw bones; teleosts have a movable premaxilla and corresponding modifications in the jaw musculature which make it possible for them to protrude their jaws outwards from the mouth. This is of great advantage, enabling them to ...
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Prehistoric Fish
The evolution of fish began about 530 million years ago during the Cambrian explosion. It was during this time that the early chordates developed the skull and the vertebral column, leading to the first craniates and vertebrates. The first fish lineages belong to the Agnatha, or jawless fish. Early examples include ''Haikouichthys''. During the late Cambrian, eel-like jawless fish called the conodonts, and small mostly armoured fish known as ostracoderms, first appeared. Most jawless fish are now extinct; but the extant lampreys may approximate ancient pre-jawed fish. Lampreys belong to the Cyclostomata, which includes the extant hagfish, and this group may have split early on from other agnathans. The earliest Gnathostomata, jawed vertebrates probably developed during the late Ordovician period. They are first represented in the fossil record from the Silurian by two groups of fish: the armoured fish known as Placodermi, placoderms, which evolved from the ostracoderms; and the ...
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