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Panoplosaurus 055
''Panoplosaurus'' is a genus of armoured dinosaur from the Late Cretaceous of Alberta, Canada. Few specimens of the genus are known, all from the middle Campanian of the Dinosaur Park Formation, roughly 76 to 75 million years ago. It was first discovered in 1917, and named in 1919 by Lawrence Lambe, named for its extensive armour, meaning "well-armoured lizard". ''Panoplosaurus'' has at times been considered the proper name for material otherwise referred to as '' Edmontonia'', complicating its phylogenetic and ecological interpretations, at one point being considered to have existed across Alberta, New Mexico and Texas, with specimens in institutions from Canada and the United States. The skull and skeleton of ''Panoplosaurus'' are similar to its relatives, but have a few significant differences, such as the lumpy form of the skull osteoderms, a completely fused scapulocoracoid, shoulder blade, and regularly shaped plates on its neck and body lacking prominent spines. It was a ...
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Campanian
The Campanian is the fifth of six ages of the Late Cretaceous Epoch on the geologic timescale of the International Commission on Stratigraphy (ICS). In chronostratigraphy, it is the fifth of six stages in the Upper Cretaceous Series. Campanian spans the time from 83.6 (± 0.2) to 72.1 (± 0.2) million years ago. It is preceded by the Santonian and it is followed by the Maastrichtian. The Campanian was an age when a worldwide sea level rise covered many coastal areas. The morphology of some of these areas has been preserved: it is an unconformity beneath a cover of marine sedimentary rocks. Etymology The Campanian was introduced in scientific literature by Henri Coquand in 1857. It is named after the French village of Champagne in the department of Charente-Maritime. The original type locality was a series of outcrop near the village of Aubeterre-sur-Dronne in the same region. Definition The base of the Campanian Stage is defined as a place in the stratigraphic column wher ...
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Skull
The skull is a bone protective cavity for the brain. The skull is composed of four types of bone i.e., cranial bones, facial bones, ear ossicles and hyoid bone. However two parts are more prominent: the cranium and the mandible. In humans, these two parts are the neurocranium and the viscerocranium ( facial skeleton) that includes the mandible as its largest bone. The skull forms the anterior-most portion of the skeleton and is a product of cephalisation—housing the brain, and several sensory structures such as the eyes, ears, nose, and mouth. In humans these sensory structures are part of the facial skeleton. Functions of the skull include protection of the brain, fixing the distance between the eyes to allow stereoscopic vision, and fixing the position of the ears to enable sound localisation of the direction and distance of sounds. In some animals, such as horned ungulates (mammals with hooves), the skull also has a defensive function by providing the mount (on the front ...
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Hadrosaurids
Hadrosaurids (), or duck-billed dinosaurs, are members of the ornithischian family Hadrosauridae. This group is known as the duck-billed dinosaurs for the flat duck-bill appearance of the bones in their snouts. The ornithopod family, which includes genera such as ''Edmontosaurus'' and ''Parasaurolophus'', was a common group of herbivores during the Late Cretaceous Period (geology), Period. Hadrosaurids are descendants of the Upper Jurassic/Lower Cretaceous iguanodontian dinosaurs and had a similar body layout. Hadrosaurs were among the most dominant herbivores during the Late Cretaceous in Asia and North America, and during the close of the Cretaceous several lineages dispersed into Europe, Africa, South America and Antarctica. Like other ornithischians, hadrosaurids had a predentary bone and a pubic bone which was positioned backwards in the pelvis. Unlike more primitive iguanodonts, the teeth of hadrosaurids are stacked into complex structures known as dental batteries, which ac ...
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Struthiosaurus
''Struthiosaurus'' (Latin ''struthio'' = ostrich + Greek ''sauros'' = lizard) is a genus of nodosaurid dinosaurs, from the Late Cretaceous period (Santonian-Maastrichtian) of Austria, Romania, France and Hungary in Europe.''Struthiosaurus'' in The Dinosaur Encyclopaedia
at Dino Russ's Lair
It was a small dinosaur, measuring in length and weighing .


History of discovery

In 1859, geologist at the ''Gute Hoffnung'' coal mine at Muthmannsdorf near

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Nodosaurus
''Nodosaurus'' (meaning "knobbed lizard") is a genus of herbivorous nodosaurid ankylosaurian dinosaur from the Late Cretaceous, the fossils of which are found exclusively in the Frontier Formation in Wyoming. Description ''Nodosaurus'' grew up to roughly long and it was an ornithischian dinosaur with bony dermal plates covering the top of its body, and it may have had spikes along its side as well. The dermal plates were arranged in bands along its body, with narrow bands over the ribs alternating with wider plates in between. These wider plates were covered in regularly arranged bony nodules, which give the animal its scientific name. In 2010 Paul estimated its length at 6 meters (20 ft) and its weight at 3.5 tonnes (3.85 short tons). It had four short legs, five-toed feet, a short neck, and a long, stiff, clubless tail. The head was narrow, with a pointed snout, powerful jaws, and small teeth. It perhaps ate soft plants, as it would have been unable to chew tough, fib ...
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Panoplosaurini
Panoplosaurini (derived from ''Panoplosaurus'', "all shield reptile") is a clade of nodosaurid ankylosaurs from the Cretaceous of North America and South America. The group is defined as the largest clade containing ''Panoplosaurus mirus'', but not '' Nodosaurus textilis'' or '' Struthiosaurus austriacus'', and was named in 2021 by Madzia and colleagues for the group found in many previous analyses, both morphological and phylogenetic. Panoplosaurini includes not only the Late Cretaceous ''Panoplosaurus'', ''Denversaurus'' and ''Edmontonia'', but also the mid Cretaceous ''Animantarx'' and ''Texasetes'', as well as ''Patagopelta''. However, in the study describing it, its authors only placed it as a nodosaurine outside Panoplosaurini. The approximately equivalent clade Panoplosaurinae, named in 1929 by Franz Nopcsa, but was not significantly used until Robert Bakker reused the name in 1988, alongside the new clades Edmontoniinae and Edmontoniidae, which were considered to unite ''Pan ...
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Animantarx
''Animantarx'' ( ; meaning 'living citadel') is a genus of nodosaurid ankylosaurian dinosaur from the Early Cretaceous, Early and Late Cretaceous of western North America. Like other nodosaurs, it would have been a slow-moving quadrupedal herbivore covered in heavy Armour (zoology), armor scutes, but without a ankylosauridae, tail club. The skull measures approximately 25 cm (10 inches) in length, suggesting the animal as a whole was no more than 3 meters (10 feet) long. Discovery and species The generic name is composed of the Latin words ''animatus'' ("living" or "animated") and ''arx'' ("fortress" or "citadel"), referring to its armored nature. In particular, the name is a reference to a comment made by paleontologist R. S. Lull about ankylosaurs, that as "an animated citadel, these animals must have been practically unassailable..." The type species is the only one known so far, and is called ''A. ramaljonesi'' after its discoverer, Ramal Jones. His wife, Carol Jones ...
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Nodosauridae
Nodosauridae is a family of ankylosaurian dinosaurs, from the Late Jurassic to the Late Cretaceous period in what is now North America, South America, Europe, and Asia. Description Nodosaurids, like their close relatives the ankylosaurids, were heavily armored dinosaurs adorned with rows of bony armor nodules and spines (osteoderms), which were covered in keratin sheaths. All nodosaurids, like other ankylosaurians, were medium-sized to large, heavily built, quadrupedal, herbivorous dinosaurs, possessing small, leaf-shaped teeth. Unlike ankylosaurids, nodosaurids lacked mace-like tail clubs, instead having flexible tail tips. Many nodosaurids had spikes projecting outward from their shoulders. One particularly well-preserved nodosaurid "mummy", known as the Suncor nodosaur (''Borealopelta markmitchelli''), preserved a nearly complete set of armor in life position, as well as the keratin covering and mineralized remains of the underlying skin, which indicate reddish dorsal pigmen ...
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Panoplosaurinae
Panoplosaurini (derived from ''Panoplosaurus'', "all shield reptile") is a clade of nodosaurid ankylosaurs from the Cretaceous of North America and South America. The group is defined as the largest clade containing ''Panoplosaurus mirus'', but not '' Nodosaurus textilis'' or '' Struthiosaurus austriacus'', and was named in 2021 by Madzia and colleagues for the group found in many previous analyses, both morphological and phylogenetic. Panoplosaurini includes not only the Late Cretaceous ''Panoplosaurus'', ''Denversaurus'' and ''Edmontonia'', but also the mid Cretaceous ''Animantarx'' and ''Texasetes'', as well as ''Patagopelta''. However, in the study describing it, its authors only placed it as a nodosaurine outside Panoplosaurini. The approximately equivalent clade Panoplosaurinae, named in 1929 by Franz Nopcsa, but was not significantly used until Robert Bakker reused the name in 1988, alongside the new clades Edmontoniinae and Edmontoniidae, which were considered to unite ''Pa ...
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Ankylosaurs
Ankylosauria is a group of herbivorous dinosaurs of the order Ornithischia. It includes the great majority of dinosaurs with armor in the form of bony osteoderms, similar to turtles. Ankylosaurs were bulky quadrupeds, with short, powerful limbs. They are known to have first appeared in the Middle Jurassic, and persisted until the end of the Cretaceous Period. The two main families of Ankylosaurs, Nodosauridae and Ankylosauridae are primarily known from the Northern Hemisphere, but the more basal Parankylosauria are known from southern Gondwana during the Cretaceous. Ankylosauria was first named by Henry Fairfield Osborn in 1923.Osborn, H. F. (1923). "Two Lower Cretaceous dinosaurs of Mongolia." ''American Museum Novitates'', 95: 1–1/ref> In the Linnaean classification system, the group is usually considered either a suborder or an infraorder. It is contained within the group Thyreophora, which also includes the stegosaurs, armored dinosaurs known for their combination of plate ...
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Ankylosaurus
''Ankylosaurus'' is a genus of armored dinosaur. Its fossils have been found in geological formations dating to the very end of the Cretaceous Period, about 68–66 million years ago, in western North America, making it among the last of the non-avian dinosaurs. It was named by Barnum Brown in 1908; it is monotypic, containing only ''A. magniventris''. The generic name means "fused" or "bent lizard", and the specific name means "great belly". A handful of specimens have been excavated to date, but a complete skeleton has not been discovered. Though other members of Ankylosauria are represented by more extensive fossil material, ''Ankylosaurus'' is often considered the archetypal member of its group, despite having some unusual features. Possibly the largest-known ankylosaurid, ''Ankylosaurus'' is estimated to have been between long and to have weighed between . It was quadrupedal, with a broad, robust body. It had a wide, low skull, with two horns pointing backward from the b ...
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Stegosaur
Stegosauria is a group of herbivorous ornithischian dinosaurs that lived during the Jurassic and early Cretaceous periods. Stegosaurian fossils have been found mostly in the Northern Hemisphere, predominantly in what is now North America, Europe, Africa, South America and Asia. Their geographical origins are unclear; the earliest unequivocal stegosaurian, ''Huayangosaurus taibaii'', lived in China. Stegosaurians were armored dinosaurs (thyreophorans). Originally, they did not differ much from more primitive members of that group, being small, low-slung, running animals protected by armored scutes. An early evolutionary innovation was the development of spikes as defensive weapons. Later species, belonging to a subgroup called the Stegosauridae, became larger, and developed long hindlimbs that no longer allowed them to run. This increased the importance of active defence by the thagomizer, which could ward off even large predators because the tail was in a higher position, pointing ...
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