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Paleoanguimorpha
Paleoanguimorpha is a clade of anguimorphs comprising Shinisauria (represented today by shinisaurids) and Goannasauria (represented today by Varanoidea which includes the families Lanthanotidae and Varanidae). Morphological studies in the past also classified helodermatids and pythonomorphs with the varanoids in the clade Platynota, while the Chinese crocodile lizard was classified as a xenosaurid. Current molecular work finds no support in these groupings and instead has found the helodermatids more related to Diploglossa in the sister clade Neoanguimorpha, while the Chinese crocodile lizard is the closet living relative to varanoids. Pythonomorphs represented by snakes today are not closely related to varanoids and are instead a sister lineage to Anguimorpha and Iguania in the clade Toxicofera. Below is the phylogeny of the paleoanguimorph lineages after Pyron ''et al.'' (2013): See also *Neoanguimorpha *Platynota Platynota is a polyphyletic group of anguimorph lizar ...
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Goannasauria
Paleoanguimorpha is a clade of anguimorphs comprising Shinisauria (represented today by shinisaurids) and Goannasauria (represented today by Varanoidea which includes the families Lanthanotidae and Varanidae). Morphological studies in the past also classified helodermatids and pythonomorphs with the varanoids in the clade Platynota, while the Chinese crocodile lizard was classified as a xenosaurid. Current molecular work finds no support in these groupings and instead has found the helodermatids more related to Diploglossa in the sister clade Neoanguimorpha, while the Chinese crocodile lizard is the closet living relative to varanoids. Pythonomorphs represented by snake Snakes are elongated, Limbless vertebrate, limbless, carnivore, carnivorous reptiles of the suborder Serpentes . Like all other Squamata, squamates, snakes are ectothermic, amniote vertebrates covered in overlapping Scale (zoology), scales. Ma ...s today are not closely related to varanoids and are ...
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Neoanguimorpha
Neoanguimorpha is a clade of anguimorphs comprising Monstersauria (represented today by helodermatids) and Diploglossa (Xenosauridae and Anguioidea). Morphological studies in the past had classified helodermatids with the varanoids in the clade Platynota, while the Chinese crocodile lizard was classified as a xenosaurid. However molecular work found no support in these groupings and instead has found the helodermatids more related to Diploglossa, while the Chinese crocodile lizard and varanoids to form the clade Paleoanguimorpha. Below is the phylogeny of the neoanguimorph lineages after Pyron ''et al.'' (2013): See also *Paleoanguimorpha Paleoanguimorpha is a clade of anguimorphs comprising Shinisauria (represented today by shinisaurids) and Goannasauria (represented today by Varanoidea which includes the families Lanthanotidae and Varanidae). Morphological studies in the past ... References Anguimorpha {{lizard-stub ...
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Anguimorpha
The Anguimorpha is a suborder of squamates. The group was named by Fürbringer in 1900 to include all autarchoglossans closer to '' Varanus'' and '' Anguis'' than ''Scincus ''Scincus'' is a genus of skinks, lizards in the family Scincidae. The genus contains four or five species, all of which are typical desert inhabitants, living in sandy and dune-like areas with a hot and dry climate. Species in the genus ''Scinc ...''. These lizards, along with iguanians and snakes, constitute the proposed "venom clade" Toxicofera of all venomous reptiles. Evolution The oldest widely accepted member of Anguimorpha is ''Dorsetisaurus'' from the Late Jurassic of Europe and North America. In 2022, the genus ''Cryptovaranoides'' was described from the latest Triassic (Rhaetian) of England. ''Cryptovaranoides'' was recovered in the study as a crown-group Anguimorph, and therefore the oldest crown group-squamate, 35 million years older than any previously known crown-group squamate. Famili ...
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Aptian
The Aptian is an age in the geologic timescale or a stage in the stratigraphic column. It is a subdivision of the Early or Lower Cretaceous Epoch or Series and encompasses the time from 121.4 ± 1.0 Ma to 113.0 ± 1.0 Ma (million years ago), approximately. The Aptian succeeds the Barremian and precedes the Albian, all part of the Lower/Early Cretaceous. The Aptian partly overlaps the upper part of the Western European Urgonian Stage. The Selli Event, also known as OAE1a, was one of two oceanic anoxic events in the Cretaceous Period, which occurred around 120 Ma and lasted approximately 1 to 1.3 million years. The Aptian extinction was a minor extinction event hypothesized to have occurred around 116 to 117 Ma.Archangelsky, Sergio.The Ticó Flora (Patagonia) and the Aptian Extinction Event" ''Acta Paleobotanica'' 41(2), 2001, pp. 115-22. Stratigraphic definitions The Aptian was named after the small city of Apt in the Provence region of France, which is also known for its cry ...
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Toxicofera
Toxicofera (Greek for "those who bear toxins") is a proposed clade of scaled reptiles (squamates) that includes the Serpentes (snakes), Anguimorpha (monitor lizards, gila monster, and alligator lizards) and Iguania (iguanas, agamas, and chameleons). Toxicofera contains about 4,600 species, (nearly 60%) of extant Squamata. It encompasses all venomous reptile species, as well as numerous related non-venomous species. There is little morphological evidence to support this grouping, however it has been recovered by all molecular analyses as of 2012. The oldest known toxicoferan is ''Cryptovaranoides'', an anguimorph from the Late Triassic of England. Cladistics Toxicofera combines the following groups from traditional classification: * Suborder Serpentes (snakes) * Suborder Iguania (iguanas, agamid lizards, chameleons, etc.) * Infraorder Anguimorpha, consisting of: ** Family Varanidae (monitor lizards) ** Family Anguidae ( alligator lizards, glass lizards, etc.) ** Family Helo ...
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Iguania
Iguania is an infraorder of squamate reptiles that includes iguanas, chameleons, agamids, and New World lizards like anoles and phrynosomatids. Using morphological features as a guide to evolutionary relationships, the Iguania are believed to form the sister group to the remainder of the Squamata, which comprise nearly 11,000 named species, roughly 2000 of which are iguanians. However, molecular information has placed Iguania well within the Squamata as sister taxa to the Anguimorpha and closely related to snakes. The order has been under debate and revisions after being classified by Charles Lewis Camp in 1923 due to difficulties finding adequate synapomorphic morphological characteristics. Most Iguanias are arboreal but there are several terrestrial groups. They usually have primitive fleshy, non-prehensile tongues, although the tongue is highly modified in chameleons. The group has a fossil record that extends back to the Early Jurassic (the oldest known member is '' B ...
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Snake
Snakes are elongated, Limbless vertebrate, limbless, carnivore, carnivorous reptiles of the suborder Serpentes . Like all other Squamata, squamates, snakes are ectothermic, amniote vertebrates covered in overlapping Scale (zoology), scales. Many species of snakes have skulls with several more joints than their lizard ancestors, enabling them to swallow prey much larger than their heads (cranial kinesis). To accommodate their narrow bodies, snakes' paired organs (such as kidneys) appear one in front of the other instead of side by side, and most have only one functional lung. Some species retain a pelvic girdle with a pair of vestigial claws on either side of the cloaca. Lizards have evolved elongate bodies without limbs or with greatly reduced limbs about twenty-five times independently via convergent evolution, leading to many lineages of legless lizards. These resemble snakes, but several common groups of legless lizards have eyelids and external ears, which snakes lack, altho ...
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Platynota
Platynota is a polyphyletic group of anguimorph lizards and thus belongs to the order Squamata of the class Reptilia. Since it was named in 1839, it has included several groups, including monitor lizards, snakes, mosasaurs, and helodermatids. Its taxonomic use still varies, as it is sometimes considered equivalent to the group Varanoidea and other times viewed as a distinct group. It is phylogenetically defined as a clade containing Varanidae (the monitor lizards). It also includes many extinct species. Description Many skeletal features support the grouping of monitor lizards, helodermatids, and several extinct species in Platynota. All platynotans have a hinged upper jaw with widely spaced teeth, each having a large base, that erupt from behind existing teeth. The teeth are plicidentine, meaning that they have highly folded layers of dentine in their centers. Many have fused frontal and nasal bones on the top of their skulls (in other lizards, these bones are separated into pa ...
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Pythonomorpha
Pythonomorpha was originally proposed by paleontologist Edward Drinker Cope (1869) as a reptilian order comprising mosasaurs, which he believed to be close relatives of Ophidia (snakes). The etymology of the term Pythonomorpha comes from the Greek ''Python'' (a monstrous snake from Greek mythology) and ''morphe'' ("form"), and refers to the generally serpentine body plan of members of the group. Cope wrote, "In the mosasauroids, we almost realize the fictions of snake-like dragons and sea-serpents, in which men have been ever prone to indulge. On account of the ophidian part of their affinities, I have called this order Pythonomorpha." Cope incorporated two families, the Clidastidae (now defunct but including only ''Clidastes'') and the Mosasauridae (including ''Macrosaurus'' Tylosaurus.html"_;"title="=Tylosaurus">''Tylosaurus''_''Mosasaurus.html" ;"title="Tylosaurus">''Tylosaurus''.html" ;"title="Tylosaurus.html" ;"title="=Tylosaurus">''Tylosaurus''">Tylosaurus.html" ;"title="=Ty ...
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Helodermatidae
The Helodermatidae or beaded lizards are a small family of lizards endemic to North America today, but formerly more widespread in the ancient past. Traditionally, the Gila monster and the Mexican beaded lizard were the only species recognized, although the latter has recently been split into several species. While the fossil record of this family may date back to as far as the Cretaceous with genera such as '' Primaderma'' and '' Paraderma'' of North America, the oldest definitive members of the Helodermatidae date to the Early Oligocene, with ''Lowesaurus matthewi'' from North America (Nebraska) and ''Euheloderma gallicum'' from Europe Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a Continent#Subcontinents, subcontinent of Eurasia ... (France). References {{Reflist Lizard families Taxa named by John Edward Gray ...
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Varanidae
The Varanidae are a family of lizards in the superfamily Varanoidea within the Anguimorpha group. The family, a group of carnivorous and frugivorous lizards, includes the living genus '' Varanus'' and a number of extinct genera more closely related to ''Varanus'' than to the earless monitor lizard (''Lanthanotus''). ''Varanus'' includes the Komodo dragon (the largest living lizard), crocodile monitor, savannah monitor, the goannas of Australia and Southeast Asia, and various other species with a similarly distinctive appearance. Their closest living relatives are the earless monitor lizard and chinese crocodile lizard. The oldest members of the family are known from the Late Cretaceous of Mongolia. Taxonomy The Varanidae were defined (using morphological characteristics) by Estes, de Queiroz and Gauthier (1988) as the clade containing the most recent common ancestor of ''Lanthanotus'' and ''Varanus'' and all of its descendants. A similar definition was formulated by Conrad ''et al ...
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Lanthanotidae
The earless monitor lizard (''Lanthanotus borneensis'') is a semiaquatic, brown lizard native to the Southeast Asian island of Borneo. It is the only living species in the family Lanthanotidae and it is related to the true monitor lizards. Taxonomy The earless monitor lizard was described in 1878 by Franz Steindachner. The genus name ''Lanthanotus'' means "hidden ear" and the species name ''borneensis'' refers to its home island of Borneo. The uniqueness of the species was recognized from the start and Steindachner placed it in its own family, Lanthanotidae. In 1899 George Albert Boulenger relegated it to the family Helodermatidae, together with the beaded lizards and gila monster. Further studies were conducted in the 1950s where it was found that although it is related to Helodermatidae, this relationship is relatively distant. The similarity is in part the result of convergent evolution and they should be recognized as separate families. Both are part of a broader Anguimorph ...
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