Overosaurus
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Overosaurus
''Overosaurus'' (meaning "Overo lizard", after the Cerro Overo locality) is an extinct genus of sauropod dinosaurs, containing only a single species, ''Overosaurus paradasorum''. This species lived approximately 86 to 84 million years ago during the latter part of the Cretaceous Period in what is now Patagonia (in southern Argentina). ''Overosaurus paradasorum'' was relatively small compared to other sauropods from Patagonia, like the saltasaurids and other aeolosaurines, estimated as approximately . It was a ground-dwelling herbivore. Discovery and naming The only known specimen of ''Overosaurus paradasorum'' was recovered at the Cerro Overo locality in Patagonia, Argentina. The specimen was collected in 2002 by researchers from the Museo Carmen Funes, the Museo Argentino Urquiza, and the Natural History Museum of Los Angeles County, in terrestrial sediments deposited during the Campanian stage of the Late Cretaceous period, approximately 84 to 78 million years ago. This speci ...
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Aeolosaurus Rionegrinus
''Aeolosaurus'' (; "Aeolus' lizard") is a genus of titanosaurian sauropod dinosaur from the Late Cretaceous Period of what is now South America. Like most sauropods, it would have been a quadrupedal herbivore with a long neck and tail. ''Aeolosaurus'' is well known for a titanosaur, as it is represented by the remains of several individuals belonging to at least two species. However, like most titanosaurs, no remains of the skull are known. The holotype of ''Aeolosaurus rionegrinus'' consists of a series of seven tail vertebrae, as well as parts of both forelimbs and the right hindlimb. It was discovered in the Angostura Colorada Formation in Argentina, which dates from the Campanian stage of the Late Cretaceous, about 83 to 74 million years ago. The species ''A. maximus'' was transferred over to the new genus ''Arrudatitan'' in 2021. Etymology This dinosaur is named after the Greek mythological figure Aeolus, Keeper of the Winds in Homer's Odyssey, because of the frequent winds ...
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Gondwanatitan Faustoi
''Gondwanatitan'' (meaning "giant from Gondwana") was a titanosaurian sauropod dinosaur. ''Gondwanatitan'' was found in Brazil, at the time part of the southern supercontinent Gondwana, in the late Cretaceous Period (70 mya). Like some other sauropods, ''Gondwanatitan'' was tall and ate tough shoots and leaves off of the tops of trees. ''G. faustois closest relative was ''Aeolosaurus''. The type species is ''Gondwanatitan faustoi'', formally described by Kellner and de Azevedo in 1999. Etymology ''Gondwanatitan'' means "Gondwana Titan", and is named after Gondwana, the supercontinent that the genus' South American range was once part of, and the Titans of classical Greek mythology. The type and only named species, ''G. faustoi'', is a patronym honoring Dr. Fausto L. de Souza Cunha, a former curator at the Museu Nacional/UFRJ who led the excavation of the type specimen. Description ''Gondwanatitan'' was a fairly small sauropod, only 7 meters (23 ft) long and weighing about 1 tonn ...
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Aeolosaurini
Aeolosaurini is an extinct clade of titanosaurian dinosaurs known from the Cretaceous period of Argentina and Brazil. Rodrigo M. Santucci and Antonio C. de Arruda-Campos (2011) in their cladistic analysis found ''Aeolosaurus'', ''Gondwanatitan'', '' Maxakalisaurus'', '' Panamericansaurus'' and '' Rinconsaurus'' to be aeolosaurids. Aeolosaurini is characterized by several synapomorphies of the caudal vertebrae, such as angled centra, elongate prezygapophyses, and neural arches shifted anteriorly relative to the centra. In life, their tails may have been strongly curved downward as a result of these traits, which may have increased the force exerted by the caudofemoralis longus muscle in retracting the hindlimb. Some aeolosaurins, such as ''Shingopana'' and ''Overosaurus'', were relatively small compared to other titanosaurs, whereas others, such as ''Aeolosaurus maximus'', were large. Phylogeny Aeolosaurini was defined by Franco-Rosas, Salgado, Rosas and Carvalho (2004) as the ste ...
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Titanosaur
Titanosaurs (or titanosaurians; members of the group Titanosauria) were a diverse group of sauropod dinosaurs, including genera from all seven continents. The titanosaurs were the last surviving group of long-necked sauropods, with taxa still thriving at the time of the extinction event at the end of the Cretaceous. This group includes some of the largest land animals known to have ever existed, such as ''Patagotitan''—estimated at long with a weight of —and the comparably-sized ''Argentinosaurus'' and ''Puertasaurus'' from the same region. The group's name alludes to the mythological Titans of ancient Greek mythology, via the type genus (now considered a '' nomen dubium)'' ''Titanosaurus''. Together with the brachiosaurids and relatives, titanosaurs make up the larger sauropod clade Titanosauriformes. Titanosaurs have long been a poorly-known group, and the relationships between titanosaur species are still not well-understood. Description Titanosauria have the largest ...
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Pitekunsaurus Macayai
''Pitekunsaurus'' is a genus of titanosaurian sauropod dinosaur from the Late Cretaceous Anacleto Formation of Neuquén, Argentina. It was described by L. Filippi and A. Garrido in 2008. The type species is ''P. macayai''. The generic name is derived from Mapudungun ''pitekun'', meaning "to discover", the epitheton honours the discoverer, oil company explorer Luis Macaya, who found the fossil in April 2004. Classification ''Pitekunsaurus'' appears to be a genus of aeolosaurid, most closely related to the genera ''Gondwanatitan'' and ''Aeolosaurus''. Its braincase closely resembles that of the aeolosaurid ''Muyelensaurus'' as well as the titanosaurs ''incertae sedis'' ''Vahiny'' and ''Jainosaurus''. It cannot be compared to the more closely related ''Aeolosaurus'' and ''Gondwanatitan ''Gondwanatitan'' (meaning "giant from Gondwana") was a titanosaurian sauropod dinosaur. ''Gondwanatitan'' was found in Brazil, at the time part of the southern supercontinent Gondwana, in th ...
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Bajo De La Carpa Formation
The Bajo de la Carpa Formation is a geologic formation of the Neuquén Basin that crops out in northern Patagonia, in the provinces of Río Negro and Neuquén, Argentina. It is the oldest of two formations belonging to the Río Colorado Subgroup within the Neuquén Group. Formerly, that subgroup was treated as a formation, and the Bajo de la Carpa Formation was known as the Bajo de la Carpa Member.Sánchez ''et al.'', 2006 At its base, this formation conformably overlies the Plottier Formation of the older Río Neuquén Subgroup, and it is in turn overlain by the Anacleto Formation, the youngest and uppermost formation of the Neuquén Group. The Bajo de la Carpa Formation can reach in thickness in some locations, and consists mainly of sandstones of various colors, all of fluvial origin, with thin layers of mudstone and siltstone in between. Geological features such as geodes, chemical nodules, impressions of raindrops, and paleosols (fossil soils) are commonly found in this ...
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Neuquén Group
The Neuquén Group is a group of geologic formations found in Argentina. Rocks in the Neuquén Group fall within the Cenomanian to early Campanian stages of the Late Cretaceous Period. It overlies the older Lohan Cura Formation and is itself overlain by the younger Allen Formation of the Malargüe Group, separated from both by unconformities, dated to 98 and 79 Ma respectively.Leanza et al., 2004, p.63 Description Deposits have been located in the provinces of Río Negro, Neuquén, and Mendoza. Although several different types of environments are represented in various sections of the Neuquén Group, the dominant regime is alluvial deposition. Many dinosaur and other fossil types have been discovered in these sediments. Subdivision There are seven formations within the Neuquén Group. These are divided into three subgroups named after major rivers in the area. In some works, the subgroups themselves are treated as formations and what usually is considered to be the formatio ...
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Sister Taxon
In phylogenetics, a sister group or sister taxon, also called an adelphotaxon, comprises the closest relative(s) of another given unit in an evolutionary tree. Definition The expression is most easily illustrated by a cladogram: Taxon A and taxon B are sister groups to each other. Taxa A and B, together with any other extant or extinct descendants of their most recent common ancestor (MRCA), form a monophyletic group, the clade AB. Clade AB and taxon C are also sister groups. Taxa A, B, and C, together with all other descendants of their MRCA form the clade ABC. The whole clade ABC is itself a subtree of a larger tree which offers yet more sister group relationships, both among the leaves and among larger, more deeply rooted clades. The tree structure shown connects through its root to the rest of the universal tree of life. In cladistic standards, taxa A, B, and C may represent specimens, species, genera, or any other taxonomic units. If A and B are at the same taxonomi ...
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Lithostrotia
Lithostrotia is a clade of derived titanosaur sauropods that lived during the Early Cretaceous and Late Cretaceous. The group was defined by Upchurch ''et al.'' in 2004 as the most recent common ancestor of ''Malawisaurus'' and ''Saltasaurus'' and all the descendants of that ancestor. Lithostrotia is derived from the Ancient Greek , meaning "inlaid with stones", referring to the fact that many known lithostrotians are preserved with osteoderms. However, osteoderms are not a distinguishing feature of the group, as the two noted by Unchurch ''et al.'' include caudal vertebrae with strongly concave front faces (procoely), although the farthest vertebrae are not procoelous. History of research In 1895, Richard Lydekker named the family Titanosauridae to summarize sauropods with procoelous (concave on the front) caudal vertebrae. The name Titanosauridae has since been widely used, and was defined by Salgado and colleagues (1997), Gonzalaz-Riga (2003), and Salgado (2003) as a node-based ...
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Phylogenetic
In biology, phylogenetics (; from Greek φυλή/ φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups of organisms. These relationships are determined by Computational phylogenetics, phylogenetic inference methods that focus on observed heritable traits, such as DNA sequences, protein amino acid sequences, or morphology. The result of such an analysis is a phylogenetic tree—a diagram containing a hypothesis of relationships that reflects the evolutionary history of a group of organisms. The tips of a phylogenetic tree can be living taxa or fossils, and represent the "end" or the present time in an evolutionary lineage. A phylogenetic diagram can be rooted or unrooted. A rooted tree diagram indicates the hypothetical common ancestor of the tree. An unrooted tree diagram (a network) makes no assumption about the ancestral line, and does ...
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Ilium (bone)
The ilium () (plural ilia) is the uppermost and largest part of the hip bone, and appears in most vertebrates including mammals and birds, but not bony fish. All reptiles have an ilium except snakes, although some snake species have a tiny bone which is considered to be an ilium. The ilium of the human is divisible into two parts, the body and the wing; the separation is indicated on the top surface by a curved line, the arcuate line, and on the external surface by the margin of the acetabulum. The name comes from the Latin (''ile'', ''ilis''), meaning "groin" or "flank". Structure The ilium consists of the body and wing. Together with the ischium and pubis, to which the ilium is connected, these form the pelvic bone, with only a faint line indicating the place of union. The body ( la, corpus) forms less than two-fifths of the acetabulum; and also forms part of the acetabular fossa. The internal surface of the body is part of the wall of the lesser pelvis and gives ...
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Caudal Vertebra
The spinal column, a defining synapomorphy shared by nearly all vertebrates,Hagfish are believed to have secondarily lost their spinal column is a moderately flexible series of vertebrae (singular vertebra), each constituting a characteristic irregular bone whose complex structure is composed primarily of bone, and secondarily of hyaline cartilage. They show variation in the proportion contributed by these two tissue types; such variations correlate on one hand with the cerebral/caudal rank (i.e., location within the backbone), and on the other with phylogenetic differences among the vertebrate taxa. The basic configuration of a vertebra varies, but the bone is its ''body'', with the central part of the body constituting the ''centrum''. The upper (closer to) and lower (further from), respectively, the cranium and its central nervous system surfaces of the vertebra body support attachment to the intervertebral discs. The posterior part of a vertebra forms a vertebral arch (in ...
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