Ophioglossidae
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Ophioglossidae
Ophioglossidae is one of the four subclasses of Polypodiopsida (ferns). This subclass consists of the ferns commonly known as whisk ferns, grape ferns, adder's-tongues and moonworts. It is equivalent to the class Psilotopsida in previous treatments, including Smith ''et al.'' (2006). The subclass contains two orders, Psilotales and Ophioglossales, whose relationship was only confirmed by molecular phylogenetic studies. Taxonomy Smith et al. (2006) carried out the first higher-level pteridophyte classification published in the molecular phylogenetic era, and considered the ferns (monilophytes), with four classes. They placed the whisk ferns and related taxa in the class Psilotopsida, with two orders. Mark W. Chase and James L. Reveal (2009) classified them as two separate subclasses, Psilotidae and Ophioglossidae, corresponding to those orders within a much broader grouping, the class Equisetopsida ''sensu lato''. Christenhusz ''et al.'', 2011, included both the Ophioglossal ...
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Fern
A fern (Polypodiopsida or Polypodiophyta ) is a member of a group of vascular plants (plants with xylem and phloem) that reproduce via spores and have neither seeds nor flowers. The polypodiophytes include all living pteridophytes except the lycopods, and differ from mosses and other bryophytes by being vascular, i.e., having specialized tissues that conduct water and nutrients and in having life cycles in which the branched sporophyte is the dominant phase. Ferns have complex leaves called megaphylls, that are more complex than the microphylls of clubmosses. Most ferns are leptosporangiate ferns. They produce coiled fiddleheads that uncoil and expand into fronds. The group includes about 10,560 known extant species. Ferns are defined here in the broad sense, being all of the Polypodiopsida, comprising both the leptosporangiate (Polypodiidae) and eusporangiate ferns, the latter group including horsetails, whisk ferns, marattioid ferns, and ophioglossoid ferns. Ferns first ...
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Polypodiopsida
A fern (Polypodiopsida or Polypodiophyta ) is a member of a group of vascular plants (plants with xylem and phloem) that reproduce via spores and have neither seeds nor flowers. The polypodiophytes include all living pteridophytes except the lycopods, and differ from mosses and other bryophytes by being vascular, i.e., having specialized tissues that conduct water and nutrients and in having life cycles in which the branched sporophyte is the dominant phase. Ferns have complex leaves called megaphylls, that are more complex than the microphylls of clubmosses. Most ferns are leptosporangiate ferns. They produce coiled fiddleheads that uncoil and expand into fronds. The group includes about 10,560 known extant species. Ferns are defined here in the broad sense, being all of the Polypodiopsida, comprising both the leptosporangiate (Polypodiidae) and eusporangiate ferns, the latter group including horsetails, whisk ferns, marattioid ferns, and ophioglossoid ferns. Ferns first ...
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Ophioglossales
Ophioglossaceae, the adder's-tongue family, is a small family of ferns. In the Pteridophyte Phylogeny Group classification of 2016 (PPG I), it is the only family in the order Ophioglossales, which together with the Psilotales is placed in the subclass Ophioglossidae. The Ophioglossidae are one of the groups traditionally known as eusporangiate ferns. Members of the family differ from other ferns in a number of ways. Many have only a single fleshy leaf at a time. Their gametophytes are subterranean and rely on fungi for energy. Description Members of Ophioglossaceae are usually terrestrial (excepting a few epiphytic species of ''Ophioglossum'') and occur in both temperate and tropical areas. They differ from the other ferns in several respects: * Many species only send up one frond or leaf-blade per year, producing only a single leaf at a time. The leaves are usually fleshy, and in temperate areas will often turn brownish or reddish during colder months. * Instead of the leptosp ...
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Pteridophyte
A pteridophyte is a vascular plant (with xylem and phloem) that disperses spores. Because pteridophytes produce neither flowers nor seeds, they are sometimes referred to as "cryptogams", meaning that their means of reproduction is hidden. Ferns, horsetails (often treated as ferns), and lycophytes (clubmosses, spikemosses, and quillworts) are all pteridophytes. However, they do not form a monophyletic group because ferns (and horsetails) are more closely related to seed plants than to lycophytes. "Pteridophyta" is thus no longer a widely accepted taxon, but the term ''pteridophyte'' remains in common parlance, as do ''pteridology'' and ''pteridologist'' as a science and its practitioner, respectively. Ferns and lycophytes share a life cycle and are often collectively treated or studied, for example by the International Association of Pteridologists and the Pteridophyte Phylogeny Group. Description Pteridophytes (ferns and lycophytes) are free-sporing vascular plants that have a lif ...
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Psilotales
Psilotaceae is a family of ferns (class Polypodiopsida) consisting of two genera, ''Psilotum'' and ''Tmesipteris'' with about a dozen species. It is the only family in the order Psilotales. Description Once thought to be descendants of early vascular plants (the Psilophyta of the Devonian period), Psilotaceae have been shown by molecular phylogenetics to be ferns (Polypodiopsida), and a sister group of the Ophioglossaceae. The family contains two genera, ''Psilotum'' and ''Tmesipteris''. The first genus, ''Psilotum'', consists of small shrubby plants of the dry tropics commonly known as "whisk ferns". The other genus, ''Tmesipteris'', is an epiphyte found in Australia, New Zealand, and New Caledonia. All members of Psilotaceae are vascular plants without any true roots. Rather, the plants are anchored by an underground system of rhizomes. The small, stem-like gametophytes of Psilotaceae are located in this rhizome system, and they aid in a plant's nutrient absorption through t ...
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Polypodiidae (plant)
The Polypodiidae, commonly called leptosporangiate ferns, formerly Leptosporangiatae, are one of four subclasses of ferns, and the largest of these, being the largest group of living ferns, including some 11,000 species worldwide. The group has also been treated as the class Pteridopsida or Polypodiopsida, although other classifications assign them a different rank. Older names for the group include Filicidae and Filicales, although at least the "water ferns" (now the Salviniales) were then treated separately. The leptosporangiate ferns are one of the four major groups of ferns, with the other three being the eusporangiate ferns comprising the marattioid ferns (Marattiidae, Marattiaceae), the horsetails (Equisetiidae, Equisetaceae), and whisk ferns and moonworts. There are approximately 8465 species of living leptosporangiate ferns, compared with about 2070 for all other ferns, totalling 10535 species of ferns. Almost a third of leptosporangiate fern species are epiphytes. These ...
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Leptosporangiate Fern
The Polypodiidae, commonly called leptosporangiate ferns, formerly Leptosporangiatae, are one of four subclasses of ferns, and the largest of these, being the largest group of living ferns, including some 11,000 species worldwide. The group has also been treated as the class Pteridopsida or Polypodiopsida, although other classifications assign them a different rank. Older names for the group include Filicidae and Filicales, although at least the "water ferns" (now the Salviniales) were then treated separately. The leptosporangiate ferns are one of the four major groups of ferns, with the other three being the eusporangiate ferns comprising the marattioid ferns (Marattiidae, Marattiaceae), the horsetails (Equisetiidae, Equisetaceae), and whisk ferns and moonworts. There are approximately 8465 species of living leptosporangiate ferns, compared with about 2070 for all other ferns, totalling 10535 species of ferns. Almost a third of leptosporangiate fern species are epiphytes. These ...
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Marattiidae
Marattiaceae is the only family of extant (living) ferns in the order Marattiales. In the Pteridophyte Phylogeny Group classification of 2016 (PPG I), Marattiales is the only order in the subclass Marattiidae. The family has six genera and about 110 species. Many are different in appearance from other ferns, having large fronds and fleshy rootstocks. Description The Marattiaceae diverged from other ferns very early in their evolutionary history and are quite different from many plants familiar to people in temperate zones. Many of them have massive, fleshy rootstocks and the largest known fronds of any fern. The Marattiaceae is one of two groups of ferns traditionally known as eusporangiate ferns, meaning that the sporangium is formed from a group of cells as opposed to a leptosporangium in which there is a single initial cell. The large fronds characteristic of the group are most readily found in the genus ''Angiopteris'', native to Australasia, Madagascar and Oceania. These fro ...
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Equisetidae
Equisetidae is one of the four subclasses of Polypodiopsida (ferns), a group of vascular plants with a fossil record going back to the Devonian. They are commonly known as horsetails. They typically grow in wet areas, with whorls of needle-like branches radiating at regular intervals from a single vertical stem. The Equisetidae were formerly regarded as a separate division of spore plants and called Equisetophyta, Arthrophyta, Calamophyta or Sphenophyta. When treated as a class, the names Equisetopsida s.s. and Sphenopsida have also been used. They are now recognized as rather close relatives of the ferns (Polypodiopsida) of which they form a specialized lineage. However, the division between the horsetails and the other ferns is so ancient that many botanists, especially paleobotanists, still regard this group as fundamentally separate at the higher level. Description The horsetails comprise photosynthesising, "segmented", hollow stems, sometimes filled with pith. At the jun ...
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Photosynthesis
Photosynthesis is a process used by plants and other organisms to convert light energy into chemical energy that, through cellular respiration, can later be released to fuel the organism's activities. Some of this chemical energy is stored in carbohydrate molecules, such as sugars and starches, which are synthesized from carbon dioxide and water – hence the name ''photosynthesis'', from the Greek ''phōs'' (), "light", and ''synthesis'' (), "putting together". Most plants, algae, and cyanobacteria perform photosynthesis; such organisms are called photoautotrophs. Photosynthesis is largely responsible for producing and maintaining the oxygen content of the Earth's atmosphere, and supplies most of the energy necessary for life on Earth. Although photosynthesis is performed differently by different species, the process always begins when energy from light is absorbed by proteins called reaction centers that contain green chlorophyll (and other colored) pigments/chromoph ...
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Sporophyte
A sporophyte () is the diploid multicellular stage in the life cycle of a plant or alga which produces asexual spores. This stage alternates with a multicellular haploid gametophyte phase. Life cycle The sporophyte develops from the zygote produced when a haploid egg cell is fertilized by a haploid sperm and each sporophyte cell therefore has a double set of chromosomes, one set from each parent. All land plants, and most multicellular algae, have life cycles in which a multicellular diploid sporophyte phase alternates with a multicellular haploid gametophyte phase. In the seed plants, the largest groups of which are the gymnosperms and flowering plants (angiosperms), the sporophyte phase is more prominent than the gametophyte, and is the familiar green plant with its roots, stem, leaves and cones or flowers. In flowering plants the gametophytes are very reduced in size, and are represented by the germinated pollen and the embryo sac. The sporophyte produces spores (hence t ...
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Eusporangiate Fern
Eusporangiate ferns are vascular spore plants, whose sporangia arise from several epidermal cells and not from a single cell as in leptosporangiate ferns. Typically these ferns have reduced root systems and sporangia that produce large amounts of spores (up to 7000 spores per sporangium in '' Christensenia'') There are four extant eusporangiate fern families, distributed among three classes. Each family is assigned to its own order. *Class Psilotopsida **Order Psilotales, family Psilotaceae – Whisk ferns (2 genera, about 17 species) **Order Ophioglossales, family Ophioglossaceae – Adder's-tongues (5 genera, about 80 species) *Class Equisetopsida **Order Equisetales, family Equisetaceae – Horsetails (1 genus, about 15 species) *Class Marattiopsida **Order Marattiales, family Marattiaceae – Marattoid ferns (6 genera, about 500 species) The following diagram shows a likely phylogenic placement of eusporangiate fern classes within the vascular plants. Cladistics While it ...
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