Ornithocheirinae
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Ornithocheirinae
Ornithocheiridae (or ornithocheirids, meaning "bird hands") is a group of pterosaurs within the suborder Pterodactyloidea. These pterosaurs were among the last to possess teeth. Members that belong to this group lived from the Early Cretaceous, Early to Late Cretaceous periods (Valanginian to Turonian stages), around 140 to 90 million years ago. Ornithocheirids are generally infamous for having an enormously controversial and very confusing taxonomy. Although agreements that these animals were related, and therefore similar to istiodactylids and pteranodontians, there is still no virtual consensus over the exact content and interrelationships of this group. Ornithocheirids were the most successful pterosaurs during their reign, they were also the largest pterosaurs before the appearance of the azhdarchids such as ''Quetzalcoatlus''. Ornithocheirids were excellent fish hunters, they used various flight techniques to catch their prey, and they are also capable of flying great dis ...
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Ferrodraco
''Ferrodraco'' ("Iron Dragon" after the ironstone the fossil was found in) is an extinct genus of anhanguerid pterosaur known from the Late Cretaceous Winton Formation of Queensland, Australia, containing the single species ''F. lentoni''. The species was named after the former mayor of Winton, Graham Thomas ‘Butch’ Lenton. It is the most complete pterosaur fossil from Australia, being known from the holotype specimen AODF 876, consisting primarily of the anterior portion of the skull and dentary, cervical vertebral centra and a partial wing. Its wingspan was estimated to be about . ''Ferrodraco'' was found to have been within the subfamily Ornithocheirinae, as sister taxon to '' Mythunga''. A recent study also recovered ''Ferrodraco'' as sister taxon to ''Mythunga'', but both placed within the family Anhangueridae, more specifically within the subfamily Tropeognathinae. ''Ferrodraco'' is also the latest surviving member of Anhangueria. Discovery and naming The holotyp ...
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Mythunga
''Mythunga'' is a genus of anhanguerid pterosaur from the late Early Cretaceous of Australia. Fossil remains of ''Mythunga'' dated back to the Albian stage of the Early Cretaceous, and the animal itself was found to be a close relative of another Australian anhanguerid called ''Ferrodraco''. Discovery and naming ''Mythunga'' is known from a partial skull, holotype QM F18896 found in April 1991 by Philip Gilmore in marine rocks of the late Albian-age Toolebuc Formation at Dunluce Station west of Hughenden, Queensland. Only the middle snout and corresponding parts of the lower jaws are known, including the rear of a left premaxilla, the lower parts of both maxillae, the rear dentaries and a right splenial. They were three-dimensionally preserved, associated in a chalk nodule. It represents a subadult individual. The fossil was prepared by Angela Hatch of the Queensland Museum, both by mechanical means and by an acid bath. The type species ''Mythunga camara'' was named and descri ...
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Coloborhynchus
''Coloborhynchus'' is a genus of pterodactyloid pterosaur belonging to the family Anhangueridae, though it has also been recovered as a member of the Ornithocheiridae in some studies. ''Coloborhynchus'' is known from the Lower Cretaceous of England (Valanginian age, 140 to 136 million years ago), and depending on which species are included, possibly the Albian and Cenomanian ages (113 to 93.9 million years ago) as well. ''Coloborhynchus'' was once thought to be the largest known toothed pterosaur, however, a specimen of the closely related '' Tropeognathus'' is now thought to have had a larger wingspan. History and classification Like many ornithocheiroid pterosaurs named during the 19th century, ''Coloborhynchus'' has a highly convoluted history of classification. Over the years numerous species have been assigned to it, and often, species have been shuffled between ''Coloborhynchus'' and related genera by various researchers. In 1874 Richard Owen, rejecting the creation by ...
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Cimoliopterus
''Cimoliopterus'' is a genus of pterosaur that lived during the Late Cretaceous in what is now England and the United States. The first known specimen, consisting of the front part of a snout including part of a crest, was discovered in the Grey Chalk Subgroup of Kent, England, and described as the new species ''Pterodactylus cuvieri'' in 1851. The specific name ''cuvieri'' honours the palaeontologist George Cuvier, whereas the genus ''Pterodactylus'' was then used for many pterosaur species that are not thought to be closely related today. It was one of the first pterosaurs to be depicted as models in Crystal Palace Park in the 1850s. The species was subsequently assigned to various other genera, including ''Ornithocheirus'' and ''Anhanguera''. In 2013, the species was moved to a new genus, as ''Cimoliopterus cuvieri''; the generic name ''Cimoliopterus'' is derived from the Greek words for "chalk" and "wing". Other specimens and species have also been assigned to or synonymi ...
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Valanginian
In the geologic timescale, the Valanginian is an age or stage of the Early or Lower Cretaceous. It spans between 139.8 ± 3.0 Ma and 132.9 ± 2.0 Ma (million years ago). The Valanginian Stage succeeds the Berriasian Stage of the Lower Cretaceous and precedes the Hauterivian Stage of the Lower Cretaceous. Stratigraphic definitions The Valanginian was first described and named by Édouard Desor in 1853. It is named after Valangin, a small town north of Neuchâtel in the Jura Mountains of Switzerland. The base of the Valanginian is at the first appearance of calpionellid species ''Calpionellites darderi'' in the stratigraphic column. A global reference section (a GSSP) had in 2009 not yet been appointed. The top of the Valanginian (the base of the Hauterivian) is at the first appearance of the ammonite genus '' Acanthodiscus''. Subdivision The Valanginian is often subdivided in Lower and Upper substages. The Upper substage begins at the first appearance of ammonite spec ...
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Late Cretaceous
The Late Cretaceous (100.5–66 Ma) is the younger of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''creta'', the Latin word for the white limestone known as chalk. The chalk of northern France and the white cliffs of south-eastern England date from the Cretaceous Period. Climate During the Late Cretaceous, the climate was warmer than present, although throughout the period a cooling trend is evident. The tropics became restricted to equatorial regions and northern latitudes experienced markedly more seasonal climatic conditions. Geography Due to plate tectonics, the Americas were gradually moving westward, causing the Atlantic Ocean to expand. The Western Interior Seaway divided North America into eastern and western halves; Appalachia and Laramidia. India maintained a northward course towards Asia. In the Southern Hemisphere, Australia an ...
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Pterodactyloidea
Pterodactyloidea (derived from the Greek words ''πτερόν'' (''pterón'', for usual ''ptéryx'') "wing", and ''δάκτυλος'' (''dáktylos'') "finger" meaning "winged finger", "wing-finger" or "finger-wing") is one of the two traditional suborders of pterosaurs ("wing lizards"), and contains the most derived members of this group of flying reptiles. They appeared during the middle Jurassic Period, and differ from the basal (though paraphyletic) rhamphorhynchoids by their short tails and long wing metacarpals (hand bones). The most advanced forms also lack teeth, and by the late Cretaceous, all known pterodactyloids were toothless. Many species had well-developed crests on the skull, a form of display taken to extremes in giant-crested forms like '' Nyctosaurus'' and '' Tupandactylus''. Pterodactyloids were the last surviving pterosaurs when the order became extinct at the end of the Cretaceous Period, together with the non-avian dinosaurs and most marine reptiles. " Pter ...
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Early Cretaceous
The Early Cretaceous (geochronology, geochronological name) or the Lower Cretaceous (chronostratigraphy, chronostratigraphic name), is the earlier or lower of the two major divisions of the Cretaceous. It is usually considered to stretch from 145 Megaannum#SI prefix multipliers, Ma to 100.5 Ma. Geology Proposals for the exact age of the Barremian-Aptian boundary ranged from 126 to 117 Ma until recently (as of 2019), but based on drillholes in Svalbard the defining Anoxic event#Cretaceous, early Aptian Oceanic Anoxic Event 1a (OAE1a) was carbon isotope dated to 123.1±0.3 Ma, limiting the possible range for the boundary to c. 122–121 Ma. There is a possible link between this anoxic event and a series of Early Cretaceous large igneous provinces (LIP). The Ontong Java Plateau, Ontong Java-Manihiki Plateau, Manihiki-Hikurangi Plateau, Hikurangi large igneous province, emplaced in the South Pacific at c. 120 Ma, is by far the largest LIP in Earth's history. T ...
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Istiodactylid
Istiodactylidae is a small family of pterosaurs. This family was named in 2001 after the type genus ''Istiodactylus'' was discovered not to be a member of the genus '' Ornithodesmus''. Systematics and distribution Remains of taxa that can be confidently assigned to Istiodactylidae have been found in the UK and China, in rocks dating from the Early Cretaceous period (Barremian to Aptian stage). Arbour and Currie (2011) described Canadian ''Gwawinapterus beardi'' as a member of Istiodactylidae living in the late Cretaceous (upper Campanian stage); however, Witton (2012) suggested the tooth replacement pattern in this animal does not match that of pterosaurs, suggesting that the species might be non-pterosaurian. Additional research suggested that the species was in fact a fish. The earliest known species might be ''Archaeoistiodactylus linglongtaensis'', from the Middle Jurassic of China; however, it also has been suggested that the holotype specimen of this species might actually ...
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Reginald Walter Hooley
Reginald Walter Hooley (5 September 1865 – 5 May 1923) was a businessman and amateur paleontologist, collecting on the Isle of Wight. He is probably best remembered for describing the dinosaur ''Iguanodon atherfieldensis'', now ''Mantellisaurus.'' Biography Reginald Hooley was born on 5 September 1865 in Southampton, the son of William Hooley, a wealthy gentleman. In 1889 R.W. Hooley began to work for ''Godrich & Petman'', wine merchants, and later in life became managing director of that firm. Living in Portswood, in 1912 he married E.E. Holden and moved to Winchester. In 1913 he was elected a member of the Winchester city council. Hooley was a member of the Hampshire Field Club & Archaeological Society at Winchester from 1890. He was one of the founders of the Isle of Wight Natural History and Archaeological Society. He was an honorary curator of the Winchester Museum between 1918 and 1923. Isle of Wight Hooley made regular visits to the Isle of Wight and found hundreds of ...
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Pteranodontia
Pteranodontia is an extinct group of ornithocheiroid pterodactyloid pterosaurs that lived during the Late Cretaceous period (Coniacian to Maastrichtian stages) of North America and Africa. They were some of the most advanced pterosaurs, and possessed highly specialized cranial crests that may have served as sexual attraction, with males having a much larger crest. Classification Pteranodontia was originally named by Marsh in 1876. In 2003, it was given a phylogenetic definition by David Unwin as the common ancestor of ''Pteranodon'' and '' Nyctosaurus'' plus all its descendants. Though Marsh had originally named this group based on the shared absence of teeth in those species, most analyses show that all of the traditional "ornithocheiroid" pterosaurs are also members of this clade. Below is a cladogram showing the phylogenetic placement of this group from Andres and Myers (2013). In 2018, Longrich, Martill, and Andres revisited the classification, and made a different phylogen ...
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Azhdarchid
Azhdarchidae (from the Persian word , , a dragon-like creature in Persian mythology) is a family of pterosaurs known primarily from the Late Cretaceous Period, though an isolated vertebra apparently from an azhdarchid is known from the Early Cretaceous as well (late Berriasian age, about 140 million years ago). Azhdarchids included some of the largest known flying animals of all time, but smaller cat-size members have also been found. Originally considered a sub-family of Pteranodontidae, Nesov (1984) named the Azhdarchinae to include the pterosaurs '' Azhdarcho'', ''Quetzalcoatlus'', and ''Titanopteryx'' (now known as '' Arambourgiania''). They were among the last known surviving members of the pterosaurs, and were a rather successful group with a worldwide distribution. By the time of the end-Cretaceous mass extinction, most pterosaur families except for the Azhdarchidae disappear from the fossil record, but recent studies indicate a wealth of pterosaurian fauna, including pter ...
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