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Oohkotokia
''Oohkotokia'' ( ) is a genus of ankylosaurid dinosaur within the subfamily Ankylosaurinae. It is known from the upper levels of the Two Medicine Formation (late Campanian stage, about 74 Ma ago) of Montana, United States. The discovery of ''Oohkotokia'' supports that Ankylosaurine dinosaurs existed and flourished continuously in Montana and/or Alberta throughout the late Campanian and early Maastrichtian stages in the Late Cretaceous period. It was a large, heavily built, quadrupedal, herbivore, that could grow up to long and weigh up to . Discovery and naming The generic name, ''Oohkotokia'', is derived from the Blackfoot animate noun "ooh’kotoka", meaning "large stone" and the Latin suffix "ia" meaning "derived from"; thus "child of stone", which is a reference to its extensive body armor. The generic name also honors the Blackfoot people, on whose land the specimen was found. The specific name, ''O. horneri'', refers to John R. Horner of the Museum of the Rockies, who ...
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Oohkotokia Skull
''Oohkotokia'' ( ) is a genus of ankylosaurid dinosaur within the subfamily Ankylosaurinae. It is known from the upper levels of the Two Medicine Formation (late Campanian stage, about 74 Ma ago) of Montana, United States. The discovery of ''Oohkotokia'' supports that Ankylosaurine dinosaurs existed and flourished continuously in Montana and/or Alberta throughout the late Campanian and early Maastrichtian stages in the Late Cretaceous period. It was a large, heavily built, quadrupedal, herbivore, that could grow up to long and weigh up to . Discovery and naming The generic name, ''Oohkotokia'', is derived from the Blackfoot language, Blackfoot animate noun "ooh’kotoka", meaning "large stone" and the Latin suffix "ia" meaning "derived from"; thus "child of stone", which is a reference to its extensive Osteoderms, body armor. The generic name also honors the Blackfoot people, on whose land the specimen was found. The specific name, ''O. horneri'', refers to John R. Horner of t ...
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Two Medicine Formation
The Two Medicine Formation is a geological formation, or rock body, in northwestern Montana and southern Alberta that was deposited between and (million years ago), during Campanian (Late Cretaceous) time. It crops out to the east of the Rocky Mountain Overthrust Belt, and the western portion (about thick) of this formation is folded and faulted while the eastern part, which thins out into the Sweetgrass Arch, is mostly undeformed plains. Below the formation are the nearshore (beach and tidal zone) deposits of the Virgelle Sandstone, and above it is the marine Bearpaw Shale. Throughout the Campanian, the Two Medicine Formation was deposited between the western shoreline of the Late Cretaceous Interior Seaway and the eastward advancing margin of the Cordilleran Overthrust Belt. The Two Medicine Formation is mostly sandstone, deposited by rivers and deltas. History of research In 1913 in paleontology, 1913, a US Geological Survey crew headed by Eugene Stebinger and a US Nationa ...
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Euoplocephalus
''Euoplocephalus'' ( ) is a genus of very large, herbivorous ankylosaurid dinosaurs, living during the Late Cretaceous of Canada. It has only one named species, ''Euoplocephalus tutus''. The first fossil of ''Euoplocephalus'' was found in 1897 in Alberta. In 1902, it was named ''Stereocephalus'', but that name had already been given to an insect, so it was changed in 1910. Later, many more ankylosaurid remains were found from the Campanian of North America and often made separate genera. In 1971, Walter Coombs concluded that they all belonged to ''Euoplocephalus'' which then would be one of the best-known dinosaurs. Recently however, experts have come to the opposite conclusion, limiting the authentic finds of ''Euoplocephalus'' to about a dozen specimens. These include a number of almost complete skeletons, so much is nevertheless known about the build of the animal. ''Euoplocephalus'' reached in length and in body mass. Its body was low-slung and very flat and wide, standin ...
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Ankylosaurine
Ankylosaurinae is a subfamily of ankylosaurid dinosaurs, existing from the Early Cretaceous about 105 million years ago until the end of the Late Cretaceous, about 66 mya. Many genera are included in the clade, such as ''Ankylosaurus'', ''Pinacosaurus'', ''Euoplocephalus'', and ''Saichania''. Features Ankylosaurines are defined as being closer relatives to ''Ankylosaurus'' than to ''Shamosaurus''. Diagnostic features of ankylosaurines include the nuchal shelf that obscures the occiput in dorsal view, and the quadrate condyle which is obscured lightly by the quadratojugal boss. Phylogeny The following cladogram is based on the 50% majority rule phylogenetic analysis In biology, phylogenetics (; from Greek φυλή/ φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups o ... of Arbour & Currie (2015): References Ankylosaur ...
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Ankylosaurid
Ankylosauridae () is a family of armored dinosaurs within Ankylosauria, and is the sister group to Nodosauridae. The oldest known Ankylosaurids date to around 122 million years ago and went extinct 66 million years ago during the Cretaceous–Paleogene extinction event. These animals were mainly herbivorous and were obligate quadrupeds, with leaf-shaped teeth and robust, scute-covered bodies. Ankylosaurids possess a distinctly domed and short snout, wedge-shaped osteoderms on their skull, scutes along their torso, and a tail club. Ankylosauridae is exclusively known from the northern hemisphere, with specimens found in western North America, Europe, and East Asia. The first discoveries within this family were of the genus ''Ankylosaurus'', by Peter Kaiser and Barnum Brown in Montana in 1906. Brown went on to name Ankylosauridae and the subfamily Ankylosaurinae in 1908. Anatomy Ankylosaurids are stout, solidly built, armoured dinosaurs. They possess accessory ossifications on c ...
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Ankylosaurinae
Ankylosaurinae is a subfamily of ankylosaurid dinosaurs, existing from the Early Cretaceous about 105 million years ago until the end of the Late Cretaceous, about 66 mya. Many genera are included in the clade, such as ''Ankylosaurus'', ''Pinacosaurus'', ''Euoplocephalus'', and ''Saichania''. Features Ankylosaurines are defined as being closer relatives to ''Ankylosaurus'' than to ''Shamosaurus''. Diagnostic features of ankylosaurines include the nuchal shelf that obscures the occiput in dorsal view, and the quadrate condyle which is obscured lightly by the quadratojugal boss. Phylogeny The following cladogram is based on the 50% majority rule phylogenetic analysis In biology, phylogenetics (; from Greek φυλή/ φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups o ... of Arbour & Currie (2015): References Ankylosaur ...
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Late Cretaceous
The Late Cretaceous (100.5–66 Ma) is the younger of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''creta'', the Latin word for the white limestone known as chalk. The chalk of northern France and the white cliffs of south-eastern England date from the Cretaceous Period. Climate During the Late Cretaceous, the climate was warmer than present, although throughout the period a cooling trend is evident. The tropics became restricted to equatorial regions and northern latitudes experienced markedly more seasonal climatic conditions. Geography Due to plate tectonics, the Americas were gradually moving westward, causing the Atlantic Ocean to expand. The Western Interior Seaway divided North America into eastern and western halves; Appalachia and Laramidia. India maintained a northward course towards Asia. In the Southern Hemisphere, Australia and Ant ...
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Osteoderm
Osteoderms are bony deposits forming scales, plates, or other structures based in the dermis. Osteoderms are found in many groups of extant and extinct reptiles and amphibians, including lizards, crocodilians, frogs, temnospondyls (extinct amphibians), various groups of dinosaurs (most notably ankylosaurs and stegosaurians), phytosaurs, aetosaurs, placodonts, and hupehsuchians (marine reptiles with possible ichthyosaur affinities). Osteoderms are uncommon in mammals, although they have occurred in many xenarthrans (armadillos and the extinct glyptodonts and mylodontid and scelidotheriid ground sloths). The heavy, bony osteoderms have evolved independently in many different lineages. The armadillo osteoderm is believed to develop in subcutaneous dermal tissues. These varied structures should be thought of as anatomical analogues, not homologues, and do not necessarily indicate monophyly. The structures are however derived from scutes, common to all classes of amniotes and ...
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Quadratojugal
The quadratojugal is a skull bone present in many vertebrates, including some living reptiles and amphibians. Anatomy and function In animals with a quadratojugal bone, it is typically found connected to the jugal (cheek) bone from the front and the squamosal bone from above. It is usually positioned at the rear lower corner of the cranium. Many modern tetrapods lack a quadratojugal bone as it has been lost or fused to other bones. Modern examples of tetrapods without a quadratojugal include salamanders, mammals, birds, and squamates (lizards and snakes). In tetrapods with a quadratojugal bone, it often forms a portion of the jaw joint. Developmentally, the quadratojugal bone is a dermal bone in the temporal series, forming the original braincase. The squamosal and quadratojugal bones together form the cheek region and may provide muscular attachments for facial muscles. In reptiles and amphibians In most modern reptiles and amphibians, the quadratojugal is a prominent, strapl ...
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Squamosal
The squamosal is a skull bone found in most reptiles, amphibians, and birds. In fishes, it is also called the pterotic bone. In most tetrapods, the squamosal and quadratojugal The quadratojugal is a skull bone present in many vertebrates, including some living reptiles and amphibians. Anatomy and function In animals with a quadratojugal bone, it is typically found connected to the jugal (cheek) bone from the front and ... bones form the cheek series of the skull. The bone forms an ancestral component of the dermal roof and is typically thin compared to other skull bones. The squamosal bone lies Anatomical terms of location, ventral to the temporal series and otic notch, and is bordered anteriorly by the Postorbital bone, postorbital. Posteriorly, the squamosal articulates with the quadrate bone, quadrate and Pterygoid bone, pterygoid bones. The squamosal is bordered anteroventrally by the jugal and ventrally by the quadratojugal. Function in reptiles In reptiles, the Quadrate ...
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Scapulocoracoid
The scapulocoracoid is the unit of the pectoral girdle that contains the coracoid and scapula. The coracoid itself is a beak-shaped bone that is commonly found in most vertebrates with a few exceptions. The scapula is commonly known as the ''shoulder blade''. The humerus is linked to the body via the scapula, and the clavicle is connected to the sternum via the scapula as well. Theria Theria (; Greek: , wild beast) is a subclass of mammals amongst the Theriiformes. Theria includes the eutherians (including the placental mammals) and the metatherians (including the marsupials) but excludes the egg-laying monotremes. Ch ...n mammals lack a scapulocoracoid. References * Vertebrates Comparative Anatomy, Function, Evolution by Kenneth V. Kardong. Page 325. Vertebrate anatomy {{Vertebrate anatomy-stub ...
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Humerus
The humerus (; ) is a long bone in the arm that runs from the shoulder to the elbow. It connects the scapula and the two bones of the lower arm, the radius and ulna, and consists of three sections. The humeral upper extremity consists of a rounded head, a narrow neck, and two short processes (tubercles, sometimes called tuberosities). The body is cylindrical in its upper portion, and more prismatic below. The lower extremity consists of 2 epicondyles, 2 processes (trochlea & capitulum), and 3 fossae (radial fossa, coronoid fossa, and olecranon fossa). As well as its true anatomical neck, the constriction below the greater and lesser tubercles of the humerus is referred to as its surgical neck due to its tendency to fracture, thus often becoming the focus of surgeons. Etymology The word "humerus" is derived from la, humerus, umerus meaning upper arm, shoulder, and is linguistically related to Gothic ''ams'' shoulder and Greek ''ōmos''. Structure Upper extremity The upper or pr ...
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