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Ankylosaurid
Ankylosauridae () is a family of armored dinosaurs within Ankylosauria, and is the sister group to Nodosauridae. The oldest known Ankylosaurids date to around 122 million years ago and went extinct 66 million years ago during the Cretaceous–Paleogene extinction event. These animals were mainly herbivorous and were obligate quadrupeds, with leaf-shaped teeth and robust, scute-covered bodies. Ankylosaurids possess a distinctly domed and short snout, wedge-shaped osteoderms on their skull, scutes along their torso, and a tail club. Ankylosauridae is exclusively known from the northern hemisphere, with specimens found in western North America, Europe, and East Asia. The first discoveries within this family were of the genus ''Ankylosaurus'', by Peter Kaiser and Barnum Brown in Montana in 1906. Brown went on to name Ankylosauridae and the subfamily Ankylosaurinae in 1908. Anatomy Ankylosaurids are stout, solidly built, armoured dinosaurs. They possess accessory ossifications on c ...
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Ankylosaurus Tail Terminology
''Ankylosaurus'' is a genus of armored dinosaur. Its fossils have been found in geological formations dating to the very end of the Cretaceous Period, about 68–66 million years ago, in western North America, making it among the last of the non-avian dinosaurs. It was named by Barnum Brown in 1908; it is monotypic, containing only ''A. magniventris''. The generic name means "fused" or "bent lizard", and the specific name means "great belly". A handful of specimens have been excavated to date, but a complete skeleton has not been discovered. Though other members of Ankylosauria are represented by more extensive fossil material, ''Ankylosaurus'' is often considered the archetypal member of its group, despite having some unusual features. Possibly the largest-known ankylosaurid, ''Ankylosaurus'' is estimated to have been between long and to have weighed between . It was quadrupedal, with a broad, robust body. It had a wide, low skull, with two horns pointing backward from the b ...
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Ankylosaurus
''Ankylosaurus'' is a genus of armored dinosaur. Its fossils have been found in geological formations dating to the very end of the Cretaceous Period, about 68–66 million years ago, in western North America, making it among the last of the non-avian dinosaurs. It was named by Barnum Brown in 1908; it is monotypic, containing only ''A. magniventris''. The generic name means "fused" or "bent lizard", and the specific name means "great belly". A handful of specimens have been excavated to date, but a complete skeleton has not been discovered. Though other members of Ankylosauria are represented by more extensive fossil material, ''Ankylosaurus'' is often considered the archetypal member of its group, despite having some unusual features. Possibly the largest-known ankylosaurid, ''Ankylosaurus'' is estimated to have been between long and to have weighed between . It was quadrupedal, with a broad, robust body. It had a wide, low skull, with two horns pointing backward from the b ...
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Ankylosaurus Magniventris
''Ankylosaurus'' is a genus of armored dinosaur. Its fossils have been found in geological formations dating to the very end of the Cretaceous Period, about 68–66 million years ago, in western North America, making it among the last of the non-avian dinosaurs. It was named by Barnum Brown in 1908; it is monotypic, containing only ''A. magniventris''. The generic name means "fused" or "bent lizard", and the specific name means "great belly". A handful of specimens have been excavated to date, but a complete skeleton has not been discovered. Though other members of Ankylosauria are represented by more extensive fossil material, ''Ankylosaurus'' is often considered the archetypal member of its group, despite having some unusual features. Possibly the largest-known ankylosaurid, ''Ankylosaurus'' is estimated to have been between long and to have weighed between . It was quadrupedal, with a broad, robust body. It had a wide, low skull, with two horns pointing backward from the b ...
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Chuanqilong
''Chuanqilong'' (meaning "legendary dragon") is a monospecific genus of basal ankylosaurid dinosaur from the Liaoning Province, China that lived during the Early Cretaceous (late Barremian to Aptian stage, 122.0 to 118.9 Ma) in what is now the Jiufotang Formation. The type and only species, ''Chuanqilong chaoyangensis'', is known from a nearly complete skeleton with a skull of a juvenile individual. It was described in 2014 by Fenglu Han, Wenjie Zheng, Dongyu Hu, Xing Xu, and Paul M. Barrett. ''Chuanqilong'' shows many similarities with '' Liaoningosaurus'' and may represent a later ontogenetic stage of the taxon. ''Chuanqilong'' was a medium-sized ankylosaur, with an estimated length of 4.5 metres (14.8 feet), although it has been suggested that it would have been larger due to the immature age of the type specimen. It had a triangular skull and a neck that was protected by bands of osteoderms known as cervical half rings. The rest of the body was covered in osteoderms and o ...
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Cedarpelta
''Cedarpelta'' is a extinct genus of basal ankylosaurid dinosaur from Utah that lived during the Late Cretaceous period (Cenomanian to lower Turonian stage, 98.2 to 93 Ma) in what is now the Mussentuchit Member of the Cedar Mountain Formation. The type and only species, ''Cedarpelta bilbeyhallorum'', is known from multiple specimens including partial skulls and postcranial material. It was named in 2001 by Kenneth Carpenter, James Kirkland, Don Burge, and John Bird. ''Cedarpelta'' has an estimated length of 7 metres (23 feet) and weight of 5 tonnes (11,023 lbs). The skull of ''Cedarpelta'' lacks extensive cranial ornamentation and is one of the only known ankylosaurs with individual skull bones that are not completely fused together. Discovery and naming The partial remains of an ankylosaur were discovered by Evan Hall and Sue Ann Bilbey at the CEM site near the Price River in Carbon County, Utah while they were visiting an excavation in the surrounding area.Carpenter, K., ...
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Scolosaurus Thronus
''Scolosaurus'' is an extinct genus of ankylosaurid dinosaurs within the subfamily Ankylosaurinae. It is known from the lower levels of the Dinosaur Park Formation and upper levels of the Oldman Formation in the Late Cretaceous (latest middle Campanian stage, about 76.5 Ma ago) of Alberta, Canada. It contains two species, ''S. cutleri'' and ''S. thronus''. The type species, ''S. cutleri'', measured up to in length and in body mass. Discovery ''Scolosaurus'' was named by Franz Nopcsa von Felső-Szilvás in 1928, based on holotype NHMUK R.5161, a nearly complete specimen that preserves the entire skeleton except for the distal end of the tail, the right forelimb, the right hindlimb, and the skull. The rare preservation of osteoderms and skin impression are also present. The fossil skeleton was discovered by William Edmund Cutler, an independent fossil collector in 1914 at Quarry 80 of the Deadlodge Canyon locality. It was collected from the bottom of the Dinosaur Park Form ...
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Aletopelta
''Aletopelta'' (; meaning 'wanderer shield') is a monospecific genus of basal ankylosaurid dinosaur from Southern California that lived during the Late Cretaceous (upper Campanian stage, 75.5 Ma) in what is now the Point Loma Formation. The type and only species, ''Aletopelta coombsi'', is known from a partial skeleton preserving osteoderms. It was originally described in 1996 by W. P. Coombs, Jr. and T.A. Deméré before being named in 2001 by Tracy Ford and James Kirkland. ''Aletopelta'' has an estimated size of 5 metres (16 feet) and weight of 2 tonnes (4,409 lbs). The holotype formed a miniature reef and was scavenged upon by invertebrates and sharks. Discovery and naming In 1987, construction work was done on the College Boulevard near Carlsbad at the Californian coast. While paleontologically surveying the work, Bradford Riney noted that a skeleton had been uncovered by a ditch dug for a sewage pipe. Within days, the specimen was secured by the San Diego Natural Histor ...
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Minmi Paravertebra
''Minmi'' is a genus of small herbivorous ankylosaurian dinosaur that lived during the early Cretaceous Period of Australia, about 133 to 120 million years ago. Discovery and species In 1964, Dr Alan Bartholomai, a collaborator of the Queensland Museum, discovered a chalkstone nodule containing an ankylosaurian skeleton in Queensland near Minmi Crossing, along the Injun Road, one kilometre south of Mack Gulley, north of Roma. In 1980, Ralph E. Molnar named and described the type species, in this case the only species known in the genus, ''Minmi paravertebra''. The generic name, at the time the shortest of a Mesozoic dinosaur, refers to Minmi Crossing. The meaning of "minmi" itself is uncertain; it refers to a large lily in the local aboriginal language but might also be derived from ''min min'', a kind of will-o'-the-wisp. The specific name refers to strange bone elements found along the vertebrae, for which Molnar coined the designation ''paravertebrae''. The holotype, ...
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Crichtonsaurus
''Crichtonsaurus'' (meaning "Crichton's lizard") is a genus of herbivorous ankylosaurid dinosaur that lived during the Late Cretaceous in what is now China. It was named after Michael Crichton, the author of the dinosaur novel ''Jurassic Park''. History The first fossils of the genus were discovered in 1999 in the Sunjiawan Formation of Xiafuxiang, near Beipiao in Liaoning Province, China. It was named and described by Dong Zhiming of the Institute of Vertebrate Paleontology and Paleoanthropology of the Chinese Academy of Sciences in 2002. The type species is ''Crichtonsaurus bohlini''. The generic name is in honor of Michael Crichton, American author whose novels include ''Jurassic Park'', ''The Andromeda Strain'' and others. The specific name honours Birger Bohlin, a Swedish paleontologist who during the 1930s took part in several paleontological expeditions to China. He described numerous Chinese ankylosaurs. As well as his work on dinosaurs and prehistoric mammals, Bohlin ...
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Zuul
''Zuul'' is a genus of herbivorous ankylosaurine dinosaur from the Campanian Judith River Formation of Montana. The type species is ''Zuul crurivastator''. It is known from a complete skull and tail, which represents the first ankylosaurin known from a complete skull and tail club, as well as the most complete ankylosaurid specimen thus far recovered from North America. The specimen also preserved ''in situ'' osteoderms, keratin, and skin remains. Discovery and naming In 2014, Theropoda Expeditions LLC was excavating a possible ''Gorgosaurus'' specimen near Havre, Montana. On 16 May, a skid-steer loader removing a -high overburden unexpectedly hit upon an ankylosaurian tail club. An almost complete ankylosaur skeleton proved to be present. As it had not been eroded on the surface, it was in pristine condition. It was found overturned, with the stomach facing upwards. The skeleton was largely articulated, but the skull and some neck vertebrae were separated from the main torso, ...
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Ankylosaurinae
Ankylosaurinae is a subfamily of ankylosaurid dinosaurs, existing from the Early Cretaceous about 105 million years ago until the end of the Late Cretaceous, about 66 mya. Many genera are included in the clade, such as ''Ankylosaurus'', ''Pinacosaurus'', ''Euoplocephalus'', and ''Saichania''. Features Ankylosaurines are defined as being closer relatives to ''Ankylosaurus'' than to ''Shamosaurus''. Diagnostic features of ankylosaurines include the nuchal shelf that obscures the occiput in dorsal view, and the quadrate condyle which is obscured lightly by the quadratojugal boss. Phylogeny The following cladogram is based on the 50% majority rule phylogenetic analysis In biology, phylogenetics (; from Greek φυλή/ φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups o ... of Arbour & Currie (2015): References Ankylosaur ...
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Gobisaurus
''Gobisaurus'' is an extinct genus of herbivorous basal ankylosaurid ankylosaur from the Upper Cretaceous (and possibly also the Lower Cretaceous) of China (''Nei Mongol Zizhiqu''). The genus is monotypic, containing only the species ''Gobisaurus domoculus''. Discovery and naming The Sino-Soviet Expeditions (1959–1960) discovered an ankylosaurian skeleton in the Gobi Desert of Inner Mongolia near Moartu, in the region of the Alashan Desert. The find was largely neglected until fossils were selected for a travelling exhibition touring the globe between 1990 and 1997, in the context of the China-Canada Dinosaur Project. The postcranial skeleton could not be located but the skull was displayed, informally labelled "Gobisaurus", at the time a ''nomen nudum''. In 2001, Matthew K. Vickaryous, Anthony P. Russell, Philip John Currie and Zhao Xijin named and described the type species ''Gobisaurus domoculus''. The generic name means "Gobi (Desert) lizard," referring to its provenan ...
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