Monstersauria
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Monstersauria
Monstersauria is a clade of anguimorph lizards, defined as all taxa more closely related to ''Heloderma'' than '' Varanus''. It includes ''Heloderma'', as well as several extinct genera, such as ''Estesia'', '' Primaderma'' and ''Gobiderma'', but it was found to be polyphyletic in the most recent and complete squamate phylogenetic analysis by Reeder ''et al.'' (2015). Classification Traditionally, Monstersauria was thought to include the modern Helodermatidae along with fossil genera such as ''Gobiderma'' and ''Estesia'' on the finding that it was a sister to Varanidae. But in more recent years, such as 2004 and 2008, more precise molecular studies have shown that the extant ''Heloderma'' is closer to Anguidae & kin than to Varanoidea. A large-scale integrated analysis on squamate phylogeny incorporating 737 characters of morphological and molecular data in 2015 analyzed the traditionally-monstersaurian fossil taxa along with the rest of the dataset, and what it found was a well- ...
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Morohasaurus
''Morohasaurus'' is an extinct genus of ''cf.'' Monstersauria, monstersaurian squamates know from a single left dentary from the Early Cretaceous (?Albian) Ohyamashimo Formation of the Sasayama Group in Tamba City, Hyogo Prefecture, Japan. The type and only species is ''Morohasaurus kamitakiensis.'' As of 2021, Ikeda ''et al.'' place this taxon ''a Cf.#Biological use, conferatur'' within Monstersauria based on general synapomorphies but without performing a phylogenetic analysis. The unique characters described in the paper to distinguish ''Morohasaurus kamitakiensis'' as a valid species include "the sinuous posteroventral rim of the dentary with a large U-shaped upper notch and small V-shaped lower notch, posteroventral corner of intramandibular septum with a weakly pointed eminence projecting posteriorly, and unicuspid, curved trenchant teeth with distinct blade-like carinae on their mesial and distal sides without grooves or serrations." If its presumed affinities are correct, ...
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Neoanguimorpha
Neoanguimorpha is a clade of anguimorphs comprising Monstersauria (represented today by helodermatids) and Diploglossa (Xenosauridae and Anguioidea). Morphological studies in the past had classified helodermatids with the varanoids in the clade Platynota, while the Chinese crocodile lizard was classified as a xenosaurid. However molecular work found no support in these groupings and instead has found the helodermatids more related to Diploglossa, while the Chinese crocodile lizard and varanoids to form the clade Paleoanguimorpha. Below is the phylogeny of the neoanguimorph lineages after Pyron ''et al.'' (2013): See also *Paleoanguimorpha Paleoanguimorpha is a clade of anguimorphs comprising Shinisauria (represented today by shinisaurids) and Goannasauria (represented today by Varanoidea which includes the families Lanthanotidae and Varanidae). Morphological studies in the past ... References Anguimorpha {{lizard-stub ...
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Anguimorpha
The Anguimorpha is a suborder of squamates. The group was named by Fürbringer in 1900 to include all autarchoglossans closer to '' Varanus'' and '' Anguis'' than ''Scincus ''Scincus'' is a genus of skinks, lizards in the family Scincidae. The genus contains four or five species, all of which are typical desert inhabitants, living in sandy and dune-like areas with a hot and dry climate. Species in the genus ''Scinc ...''. These lizards, along with iguanians and snakes, constitute the proposed "venom clade" Toxicofera of all venomous reptiles. Evolution The oldest widely accepted member of Anguimorpha is ''Dorsetisaurus'' from the Late Jurassic of Europe and North America. In 2022, the genus ''Cryptovaranoides'' was described from the latest Triassic (Rhaetian) of England. ''Cryptovaranoides'' was recovered in the study as a crown-group Anguimorph, and therefore the oldest crown group-squamate, 35 million years older than any previously known crown-group squamate. Famili ...
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Estesia
''Estesia'' (in honour of Richard Estes) is an extinct genus of Late Cretaceous anguimorph lizard found in the Gobi Desert in Mongolia. It was discovered in June 1990 by a joint expedition made up of Mongolian and American palaeontologists, and described in 1992 by Mark Norell, Malcolm McKenna and Michael Novacek. This animal is of interest to palaeontologists, not only because it is close to the lineage of modern Gila monsters (''Heloderma''), but also because its dentition shows evidence that it was venomous. The type species is ''E. mongoliensis'', after Mongolia, where it was found. Material Material for ''Estesia'' has been collected in various localities in the Gobi Desert, including the Barun Goyot Formation, the Djadochta Formation and Ukhaa Tolgod. *IGM M3/14 (holotype): A well-preserved skull with mandible, from Lizard Hill, Khulsan (Barun Goyot Formation) *IGM 3/15: Partial skeleton with associated braincase, from Khermeen Tsav (Barun Goyot Formation), discovered ...
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Asprosaurus
''Asprosaurus'' is an extinct genus of anguimorph lizard from the Late Cretaceous of South Korea. Named in 2015 from the Seonso Conglomerate Formation, the type species ''Asprosaurus bibongriensis'' is the first Mesozoic lizard to have been discovered on the Korean peninsula. Because ''Asprosaurus'' is known only from fragmentary material, its relationships with other lizards are uncertain. However, features of the lower jaw suggest that it may be a member of a clade (evolutionary grouping) called Monstersauria, which includes the living Gila monster. Size ''Asprosaurus'' is among the largest Late Cretaceous terrestrial lizards from Asia described to date, with an estimated skull length of , only rivalled by '' Chianghsia'' from China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones ...
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Huehuecuetzpalli
''Huehuecuetzpalli mixtecus'' is an extinct lizard from the Early Cretaceous (specifically the middle to late Aptian stage) Tlayúa Formation in Tepexi de Rodríguez (municipality), Tepexi de Rodríguez, Central Mexico. Although it is not the oldest known lizard, ''Huehuecuetzpalli'' may be amongst the most basal (phylogenetics), basal members of Squamata (the group that includes lizards and snakes), making it an important taxon in understanding the origins of squamates. The generic name comes from the Nahuatl words ''huehuetl'' ("the ancient") and ''cuetzpalli'' ("lizard"), while the specific name refers to the La Mixteca region. Description Unique characteristics (autapomorphies) of ''Huehuecuetzpalli'' include a long pair of premaxilla bones at the tip of the upper jaw that contributes to an elongated snout and the apparent retraction of the nostril, external nares or nostril openings. At the top of the skull, a small rounded postfrontal and a hole called the Pineal foramen, ...
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Anguidae
Anguidae refers to a large and diverse family of lizards native to the Northern Hemisphere. Common characteristics of this group include a reduced supratemporal arch, striations on the medial faces of tooth crowns, osteoderms, and a lateral fold in the skin of most taxa. The group includes the slowworms, glass lizards, and alligator lizards, among others. The family is divided into two subfamilies (Anguinae and Gerrhonotinae), and contains about 87 species in 8 genera. Morphology and reproduction Anguids have hard osteoderms beneath their scales giving them an armored appearance. Many of the species have reduced or absent limbs, giving them a snake-like appearance, while others are fully limbed. Body type varies among species, with sizes ranging from 10 cm to 1.5 m. The group includes oviparous and viviparous species, both of which can be observed in a single genus at times. Feeding and habitat These lizards are known carnivorous or insectivorous foragers, feeding p ...
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Varanoidea
Varanoidea is a superfamily of lizards, including the well-known family Varanidae (the monitors and goannas). Also included in the Varanoidea are the Lanthanotidae (earless monitor lizards), and the extinct Palaeovaranidae. Throughout their long evolutionary history, varanoids have exhibited great diversity, both in habitat and form. This superfamily includes the largest-known terrestrial lizard, Megalania (5–6 meters), and the largest extant lizard, the Komodo dragon (''Varanus komodoensis'', 3+ meters). Evolution Either synonymous with, or a subgroup of, the group Platynota, the varanoids first appear in the fossil record in the latter part of Early Cretaceous, but possible varanoid ancestors have been traced back as far as Early Jurassic times. Among the earliest known varanoids are the monitor-like necrosaurids '' Palaeosaniwa canadensis'' from the Campanian (roughly 71-82 mya) of North America and '' Estesia mongoliensis'' and '' Telmasaurus grangeri'', both from the Cam ...
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Squamata
Squamata (, Latin ''squamatus'', 'scaly, having scales') is the largest order of reptiles, comprising lizards, snakes, and amphisbaenians (worm lizards), which are collectively known as squamates or scaled reptiles. With over 10,900 species, it is also the second-largest order of extant (living) vertebrates, after the perciform fish. Members of the order are distinguished by their skins, which bear horny scales or shields, and must periodically engage in molting. They also possess movable quadrate bones, making possible movement of the upper jaw relative to the neurocranium. This is particularly visible in snakes, which are able to open their mouths very wide to accommodate comparatively large prey. Squamata is the most variably sized order of reptiles, ranging from the dwarf gecko (''Sphaerodactylus ariasae'') to the Reticulated python (''Malayopython reticulatus'') and the now-extinct mosasaurs, which reached lengths over . Among other reptiles, squamates are most close ...
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Gila Monster
The Gila monster (''Heloderma suspectum'', ) is a species of venomous lizard native to the Southwestern United States and the northwestern Mexican state of Sonora. It is a heavy, typically slow-moving reptile, up to long, and it is the only venomous lizard native to the United States. Its venomous close relatives, the four beaded lizards (all former subspecies of ''Heloderma horridum'') inhabit Mexico and Guatemala. The Gila monster is sluggish in nature, so it is not generally dangerous and very rarely poses a real threat to humans. However, it has a fearsome reputation and is sometimes killed in spite of the species being protected by state law in Arizona. History The name "Gila" refers to the Gila River Basin in the U.S. states of Arizona and New Mexico, where the Gila monster was once plentiful. ''Heloderma'' means "studded skin", from the Ancient Greek words (), "the head of a nail or stud", and (), "skin". ''Suspectum'' comes from the describer, paleontologist Edwa ...
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