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Metachronal Swimming
A metachronal swimming or metachronal rowing is the swimming technique used by animals with multiple pairs of swimming legs. In this technique, appendages are sequentially stroked in a back-to-front wave moving along the animal’s body. In literature, while metachronal rhythm or metachronal wave usually refer to the movement of cilia; metachronal coordination, metachronal beating, metachronal swimming or metachronal rowing usually refer to the leg movement of arthropods, such as mantis shrimp, copepods, antarctic krill etc. though all of them refer to the similar locomotion pattern. Metachronous indicates something not functioning or occurring synchronously, or occurring or starting at different times. This word is derived from Greek meta- μετά- meaning, occurring later than or in succession to : after, and -chronous -Χρόνος meaning, of (such) a time or period. Swimming legs should coordinate to avoid interference among appendage pairs. To accomplish this challenge, almo ...
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Appendage
An appendage (or outgrowth) is an external body part, or natural prolongation, that protrudes from an organism's body. In arthropods, an appendage refers to any of the homologous body parts that may extend from a body segment, including antennae, mouthparts (including mandibles, maxillae and maxillipeds), gills, locomotor legs ( pereiopods for walking, and pleopods for swimming), sexual organs (gonopods), and parts of the tail (uropods). Typically, each body segment carries one pair of appendages. An appendage which is modified to assist in feeding is known as a maxilliped or gnathopod. In vertebrates, an appendage can refer to a locomotor part such as a tail, fins on a fish, limbs (legs, flippers or wings) on a tetrapod; exposed sex organ; defensive parts such as horns and antlers; or sensory organs such as auricles, proboscis ( trunk and snout) and barbels. Appendages may become ''uniramous'', as in insects and centipedes, where each appendage comprises a single ...
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Metachronal Rhythm
A metachronal rhythm or metachronal wave refers to wavy movements produced by the sequential action (as opposed to synchronized) of structures such as cilia, segments of worms, or legs. These movements produce the appearance of a travelling wave. A Mexican wave is a large scale example of a metachronal wave. This pattern is found widely in nature such as on the cilia of many aquatic organisms such as ctenophores, molluscs, ciliates as well as on the epithelial surfaces of many body organs. Individual cilia, when part of a metachronal wave being used for protist locomotion, individually beat in a pattern similar to the planar stroke of a flagellum. The difference is that the recovery stroke is at 90 degrees to the power stroke, so that the cilia avoid hitting each other. Metachronal rhythms may be seen in the coordinated movements of the legs of millipedes and other multi-legged land invertebrates, as well as in the coordinated movements of social insects. Such metachronal mo ...
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Cilium
The cilium, plural cilia (), is a membrane-bound organelle found on most types of eukaryotic cell, and certain microorganisms known as ciliates. Cilia are absent in bacteria and archaea. The cilium has the shape of a slender threadlike projection that extends from the surface of the much larger cell body. Eukaryotic flagella found on sperm cells and many protozoans have a similar structure to motile cilia that enables swimming through liquids; they are longer than cilia and have a different undulating motion. There are two major classes of cilia: ''motile'' and ''non-motile'' cilia, each with a subtype, giving four types in all. A cell will typically have one primary cilium or many motile cilia. The structure of the cilium core called the axoneme determines the cilium class. Most motile cilia have a central pair of single microtubules surrounded by nine pairs of double microtubules called a 9+2 axoneme. Most non-motile cilia have a 9+0 axoneme that lacks the central pair o ...
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Arthropod
Arthropods (, (gen. ποδός)) are invertebrate animals with an exoskeleton, a Segmentation (biology), segmented body, and paired jointed appendages. Arthropods form the phylum Arthropoda. They are distinguished by their jointed limbs and Arthropod cuticle, cuticle made of chitin, often Mineralization (biology), mineralised with calcium carbonate. The arthropod body plan consists of segments, each with a pair of appendages. Arthropods are bilaterally symmetrical and their body possesses an exoskeleton, external skeleton. In order to keep growing, they must go through stages of moulting, a process by which they shed their exoskeleton to reveal a new one. Some species have wings. They are an extremely diverse group, with up to 10 million species. The haemocoel, an arthropod's internal cavity, through which its haemolymph – analogue of blood – circulates, accommodates its interior Organ (anatomy), organs; it has an open circulatory system. Like their exteriors, the internal or ...
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Mantis Shrimp
Mantis shrimp, or stomatopods, are carnivorous marine crustaceans of the order Stomatopoda (). Stomatopods branched off from other members of the class Malacostraca around 340 million years ago. Mantis shrimp typically grow to around in length, while a few can reach up to . A mantis shrimp's carapace (the hard, thick shell that covers crustaceans and some other species) covers only the rear part of the head and the first four segments of the thorax. Varieties range in colour from shades of brown to vivid colours, with more than 450 species of mantis shrimp known. They are among the most important predators in many shallow, tropical and subtropical marine habitats. However, despite being common, they are poorly understood, as many species spend most of their lives sheltering in burrows and holes. Called "sea locusts" by ancient Assyrians, "prawn killers" in Australia, and now sometimes referred to as "thumb splitters"—because of the animal's ability to inflict painful woun ...
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Copepod
Copepods (; meaning "oar-feet") are a group of small crustaceans found in nearly every freshwater and saltwater habitat (ecology), habitat. Some species are planktonic (inhabiting sea waters), some are benthos, benthic (living on the ocean floor), a number of species have parasitic phases, and some continental species may live in limnoterrestrial habitats and other wet terrestrial places, such as swamps, under leaf fall in wet forests, bogs, springs, ephemeral ponds, and puddles, damp moss, or water-filled recesses (phytotelmata) of plants such as bromeliads and pitcher plants. Many live underground in marine and freshwater caves, sinkholes, or stream beds. Copepods are sometimes used as Ecological indicator, biodiversity indicators. As with other crustaceans, copepods have a larval form. For copepods, the egg hatches into a Crustacean larvae#Nauplius, nauplius form, with a head and a tail but no true thorax or abdomen. The larva molts several times until it resembles the adult an ...
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Antarctic Krill
Antarctic krill (''Euphausia superba'') is a species of krill found in the Antarctic waters of the Southern Ocean. It is a small, swimming crustacean that lives in large schools, called swarms, sometimes reaching densities of 10,000–30,000 individual animals per cubic metre. It feeds directly on minute phytoplankton, thereby using the primary production energy that the phytoplankton originally derived from the sun in order to sustain their pelagic (open ocean) life cycle. It grows to a length of , weighs up to , and can live for up to six years. It is a key species in the Antarctic ecosystem and in terms of biomass, is one of the most abundant animal species on the planet (approximately 500 million tons, corresponding to 300 to 400 trillion individuals). Life cycle The main spawning season of Antarctic krill is from January to March, both above the continental shelf and also in the upper region of deep sea oceanic areas. In the typical way of all krill, the male attaches a ...
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Animal Locomotion
Animal locomotion, in ethology, is any of a variety of methods that animal (biology), animals use to move from one place to another. Some modes of locomotion are (initially) self-propelled, e.g., running, swimming, jumping, flying, hopping, soaring and gliding. There are also many animal species that depend on their environment for transportation, a type of mobility called Passive locomotion in animals, passive locomotion, e.g., sailing (some jellyfish), Ballooning (spider), kiting (spiders), rolling (some beetles and spiders) or riding other animals (phoresis (biology), phoresis). Animals move for a variety of reasons, such as to find Foraging, food, a Mating system, mate, a suitable microhabitat, or to Escape response, escape predators. For many animals, the ability to move is essential for survival and, as a result, natural selection has shaped the locomotion methods and mechanisms used by moving organisms. For example, Animal migration, migratory animals that travel vast dista ...
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Euphausia Superba
Antarctic krill (''Euphausia superba'') is a species of krill found in the Antarctic waters of the Southern Ocean. It is a small, swimming crustacean that lives in large schools, called swarms, sometimes reaching densities of 10,000–30,000 individual animals per cubic metre. It feeds directly on minute phytoplankton, thereby using the primary production energy that the phytoplankton originally derived from the sun in order to sustain their pelagic (open ocean) life cycle. It grows to a length of , weighs up to , and can live for up to six years. It is a key species in the Antarctic ecosystem and in terms of biomass, is one of the most abundant animal species on the planet (approximately 500 million tons, corresponding to 300 to 400 trillion individuals). Life cycle The main spawning season of Antarctic krill is from January to March, both above the continental shelf and also in the upper region of deep sea oceanic areas. In the typical way of all krill, the male attaches a s ...
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Biomechanics
Biomechanics is the study of the structure, function and motion of the mechanical aspects of biological systems, at any level from whole organisms to organs, cells and cell organelles, using the methods of mechanics. Biomechanics is a branch of biophysics. In 2022, computational mechanics goes far beyond pure mechanics, and involves other physical actions: chemistry, heat and mass transfer, electric and magnetic stimuli and many others. Etymology The word "biomechanics" (1899) and the related "biomechanical" (1856) come from the Ancient Greek βίος ''bios'' "life" and μηχανική, ''mēchanikē'' "mechanics", to refer to the study of the mechanical principles of living organisms, particularly their movement and structure. Subfields Biofluid mechanics Biological fluid mechanics, or biofluid mechanics, is the study of both gas and liquid fluid flows in or around biological organisms. An often studied liquid biofluid problem is that of blood flow in the human card ...
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Animal
Animals are multicellular, eukaryotic organisms in the Kingdom (biology), biological kingdom Animalia. With few exceptions, animals Heterotroph, consume organic material, Cellular respiration#Aerobic respiration, breathe oxygen, are Motility, able to move, can Sexual reproduction, reproduce sexually, and go through an ontogenetic stage in which their body consists of a hollow sphere of Cell (biology), cells, the blastula, during Embryogenesis, embryonic development. Over 1.5 million Extant taxon, living animal species have been Species description, described—of which around 1 million are Insecta, insects—but it has been estimated there are over 7 million animal species in total. Animals range in length from to . They have Ecology, complex interactions with each other and their environments, forming intricate food webs. The scientific study of animals is known as zoology. Most living animal species are in Bilateria, a clade whose members have a Symmetry in biology#Bilate ...
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Antenna (biology)
Antennae ( antenna), sometimes referred to as "feelers", are paired appendages used for sensing in arthropods. Antennae are connected to the first one or two segments of the arthropod head. They vary widely in form but are always made of one or more jointed segments. While they are typically sensory organs, the exact nature of what they sense and how they sense it is not the same in all groups. Functions may variously include sensing touch, air motion, heat, vibration (sound), and especially smell or taste. Antennae are sometimes modified for other purposes, such as mating, brooding, swimming, and even anchoring the arthropod to a substrate. Larval arthropods have antennae that differ from those of the adult. Many crustaceans, for example, have free-swimming larvae that use their antennae for swimming. Antennae can also locate other group members if the insect lives in a group, like the ant. The common ancestor of all arthropods likely had one pair of uniramous (unbranched ...
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