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Mandibulate
Mandibulata, termed "mandibulates", is a clade of arthropods that comprises the extant subphyla Myriapoda (millipedes and others), Crustacea and Hexapoda (insects and others). Mandibulata is currently believed to be the sister group of the clade Arachnomorpha, which comprises the rest of arthropods (Chelicerata and Trilobita). The mandibulates constitute the largest and most varied arthropod group. The name "Mandibulata" refers to the mandibles or jaws, which are the characterizing feature of its member arthropods. Molecular phylogenetic studies suggest that the living arthropods are related as shown in the cladogram below. Crustaceans do not form a monophyletic group as insects and other hexapods have evolved from within them. See also *Atelocerata * Marrellomorpha * Myriochelata *Pancrustacea *Crustaceomorpha Crustaceomorpha is a proposed clade of arthropods that includes crustaceans and numerous extinct groups. Synapomorphies for the clade are that the larval antenn ...
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Hymenocarina
Hymenocarina is an order of extinct arthropods known from the Cambrian. They possess bivalved carapaces, typically with exposed posteriors. Members of the group are morphologically diverse and had a variety of ecologies, including as filter feeders and as predators. Recent research has generally considered them to be stem or crown group members of Mandibulata, due the presence of mandibles in at least some species. Taxonomy Hymenocarines are characterized by the combination of following characters: bivalved, convex carapace covering cephalothoracic region; cephalothorax bearing multisegmented antennules and rounded mandibles, alongside post-maxillular limbs with spiny, subdivided basis and endopods with well-developed terminal claws; absence of appendages between antennules and mandibles; median sclerite and lobate protrusions located between compound eyes; posterior tagma (abdomen) with ring-like segments and terminated by a pair of well-developed caudal rami. Based on th ...
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Mandible (arthropod Mouthpart)
250px, The mandibles of a bull ant The mandible (from la, mandibula or mandĭbŭ-lum, a jaw) of an arthropod is a pair of mouthparts used either for biting or cutting and holding food. Mandibles are often simply called jaws. Mandibles are present in the extant subphyla Myriapoda (millipedes and others), Crustacea and Hexapoda (insects etc.). These groups make up the clade Mandibulata, which is currently believed to be the sister group to the rest of arthropods, the clade Arachnomorpha (Chelicerata and Trilobita). Unlike the chelicerae of arachnids, mandibles can often be used to chew food. Mandibulates also differ by having antennae, and also by having three distinct body regions: head, thorax and abdomen. (The cephalothorax (or prosoma) of chelicerates is a fusion of head and thorax.) Insects Insect mandibles are as diverse in form as their food. For instance, grasshoppers and many other plant-eating insects have sharp-edged mandibles that move side to side. Most ...
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Bull Ant
''Myrmecia'' is a genus of ants first established by Danish zoologist Johan Christian Fabricius in 1804. The genus is a member of the subfamily Myrmeciinae of the family Formicidae. ''Myrmecia'' is a large genus of ants, comprising at least 93 species that are found throughout Australia and its coastal islands, while a single species is only known from New Caledonia. One species has been introduced out of its natural distribution and was found in New Zealand in 1940, but the ant was last seen in 1981. These ants are commonly known as bull ants, bulldog ants or jack jumper ants, and are also associated with many other common names. They are characterized by their extreme aggressiveness, ferocity, and painful stings. Some species are known for the jumping behavior they exhibit when agitated. Species of this genus are also characterized by their elongated mandibles and large compound eyes that provide excellent vision. They vary in colour and size, ranging from . Although work ...
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Crustacean
Crustaceans (Crustacea, ) form a large, diverse arthropod taxon which includes such animals as decapods, seed shrimp, branchiopods, fish lice, krill, remipedes, isopods, barnacles, copepods, amphipods and mantis shrimp. The crustacean group can be treated as a subphylum under the clade Mandibulata. It is now well accepted that the hexapods emerged deep in the Crustacean group, with the completed group referred to as Pancrustacea. Some crustaceans ( Remipedia, Cephalocarida, Branchiopoda) are more closely related to insects and the other hexapods than they are to certain other crustaceans. The 67,000 described species range in size from '' Stygotantulus stocki'' at , to the Japanese spider crab with a leg span of up to and a mass of . Like other arthropods, crustaceans have an exoskeleton, which they moult to grow. They are distinguished from other groups of arthropods, such as insects, myriapods and chelicerates, by the possession of biramous (two-parted) limbs, and by ...
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Crustaceomorpha
Crustaceomorpha is a proposed clade of arthropods that includes crustaceans and numerous extinct groups. Synapomorphies for the clade are that the larval antenna is a feeding or locomotory organ, and there are six endopodal podomeres in post-antennal limbs. Extinct groups included in Crustaceomorpha vary considerably. It includes Agnostida (usually treated under Trilobita), Waptiida, Isoxyida, Phosphatocopida, and Bradoriida, among others. These primitive crustaceomorphs are grouped under Pseudocrustacea. The validity of Crustaceomorpha is controversial. It has more support among paleontologists who consider it a sister group to Arachnomorpha, which includes trilobites and chelicerates (see cladogram below). Both are grouped under Schizoramia, a clade of arthropods with biramous appendages. In contrast, neontologists tend to support the Mandibulata clade, which groups members of Crustacea together with Hexapoda and Myriapoda. See also * Pancrustacea *Mandibulata M ...
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Myriochelata
The Myriochelata or Paradoxopoda, is a proposed grouping of arthropods comprising the Myriapoda (including millipedes and centipedes) and Chelicerata (including spiders and scorpions). If this proposition holds true, the Myriochelata are the sister clade to the Pancrustacea (also ''Tetraconata''), comprising classic crustaceans and hexapods. The evidence for this relationship between myriapods and chelicerates derives from comparisons of nuclear ribosomal RNA genes, mitochondrial ribosomal RNA genes, and protein-coding genes. More recent molecular studies, however, favour grouping the myriapods with the Pancrustacea, rather than with the chelicerates, to make up the clade Mandibulata Mandibulata, termed "mandibulates", is a clade of arthropods that comprises the extant subphyla Myriapoda (millipedes and others), Crustacea and Hexapoda (insects and others). Mandibulata is currently believed to be the sister group of the clade ....*Omar Rota-Stabelli, Lahcen Campbell, Henner Brin ...
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Marrellomorph
Marrellomorpha are an extinct group of arthropods known from the Cambrian to the Early Devonian. They lacked mineralised hard parts, so are only known from areas of exceptional preservation, limiting their fossil distribution. The best known member is ''Marrella'', with thousands of specimens found in the Cambrian aged Burgess Shale of Canada. The group is divided up into two major orders, Marrellida and Acercostraca. Marrellida is recognised by the possession of head shields with two or three pairs of elongate spine-like projections, and three pairs of uniramous appendages on the cephalon, while Acercostraca generally have large ovoid carapaces that cover the entire upper half of the body, and five pairs of uniramous cephalic appendages. Both groups have unbranched antennules and a segmented trunk with biramous appendages. Recent research has suggested the previously enigmatic Cambrian trliobite-like arthropods '' Skania'' and '' Primicaris'' may belong to this group. Their ...
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Atelocerata
Atelocerata is a proposed clade of arthropods that includes Hexapoda ( insects and a few related taxa) and Myriapoda ( millipedes, centipedes, and similar taxa), but excludes Crustacea (such as shrimp and lobsters) and Chelicerata (such as spiders and horseshoe crabs). The name is currently used interchangeably with Tracheata. or Uniramia '' sensu stricto''. It is an extensive division of arthropods comprising all those that breathe by tracheae The trachea, also known as the windpipe, is a cartilaginous tube that connects the larynx to the bronchi of the lungs, allowing the passage of air, and so is present in almost all air-breathing animals with lungs. The trachea extends from the la ..., as distinguished from Crustacea, which breathe by means of gills. The name Tracheata is an older term, originally proposed by Ernst Haeckel in 1866 as a grouping of arachnids, myriapods and insects. Tracheata was redefined to exclude arachnids by Pocock in 1893.(reviewed in ...
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Arthropoda
Arthropods (, (gen. ποδός)) are invertebrate animals with an exoskeleton, a segmented body, and paired jointed appendages. Arthropods form the phylum Arthropoda. They are distinguished by their jointed limbs and cuticle made of chitin, often mineralised with calcium carbonate. The arthropod body plan consists of segments, each with a pair of appendages. Arthropods are bilaterally symmetrical and their body possesses an external skeleton. In order to keep growing, they must go through stages of moulting, a process by which they shed their exoskeleton to reveal a new one. Some species have wings. They are an extremely diverse group, with up to 10 million species. The haemocoel, an arthropod's internal cavity, through which its haemolymph – analogue of blood – circulates, accommodates its interior organs; it has an open circulatory system. Like their exteriors, the internal organs of arthropods are generally built of repeated segments. Their nervous system is "lad ...
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Monophyly
In cladistics for a group of organisms, monophyly is the condition of being a clade—that is, a group of taxa composed only of a common ancestor (or more precisely an ancestral population) and all of its lineal descendants. Monophyletic groups are typically characterised by shared derived characteristics ( synapomorphies), which distinguish organisms in the clade from other organisms. An equivalent term is holophyly. The word "mono-phyly" means "one-tribe" in Greek. Monophyly is contrasted with paraphyly and polyphyly as shown in the second diagram. A ''paraphyletic group'' consists of all of the descendants of a common ancestor minus one or more monophyletic groups. A '' polyphyletic group'' is characterized by convergent features or habits of scientific interest (for example, night-active primates, fruit trees, aquatic insects). The features by which a polyphyletic group is differentiated from others are not inherited from a common ancestor. These definitions have tak ...
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Cladogram
A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to descendants, nor does it show how much they have changed, so many differing evolutionary trees can be consistent with the same cladogram. A cladogram uses lines that branch off in different directions ending at a clade, a group of organisms with a last common ancestor. There are many shapes of cladograms but they all have lines that branch off from other lines. The lines can be traced back to where they branch off. These branching off points represent a hypothetical ancestor (not an actual entity) which can be inferred to exhibit the traits shared among the terminal taxa above it. This hypothetical ancestor might then provide clues about the order of evolution of various features, adaptation, and other evolutionary narratives about an ...
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Molecular Phylogenetics
Molecular phylogenetics () is the branch of phylogeny that analyzes genetic, hereditary molecular differences, predominantly in DNA sequences, to gain information on an organism's evolutionary relationships. From these analyses, it is possible to determine the processes by which diversity among species has been achieved. The result of a molecular phylogenetic analysis is expressed in a phylogenetic tree. Molecular phylogenetics is one aspect of molecular systematics, a broader term that also includes the use of molecular data in taxonomy and biogeography. Molecular phylogenetics and molecular evolution correlate. Molecular evolution is the process of selective changes (mutations) at a molecular level (genes, proteins, etc.) throughout various branches in the tree of life (evolution). Molecular phylogenetics makes inferences of the evolutionary relationships that arise due to molecular evolution and results in the construction of a phylogenetic tree. History The theoretical f ...
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