Lirainosaurinae
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Lirainosaurinae
Lirainosaurinae is a subfamily of lithostrotian titanosaur sauropods from the Late Cretaceous of France, Spain, and Romania. Systematics Lirainosaurinae was defined by Díez Díaz ''et al.'' (2018) as comprising "''Lirainosaurus'', ''Ampelosaurus'', their common ancestor, and all of its descendants", and includes the two definitional taxa as well as ''Atsinganosaurus'', '' Lohuecotitan'', and ''Paludititan''. The five genera are known from Europe and all within the timespan of the late Campanian (''Lirainosaurus'') to early Maastrichtian (''Ampelosaurus'' and ''Atsinganosaurus''). Lirainosaurinae was recovered by Diez ''et al.'' (2018) as phylogenetically intermediate between the clades traditionally considered Saltasauridae and a clade containing taxa normally found in Aeolosaurini and Lognkosauria Lognkosauria is a clade of giant long-necked sauropod dinosaurs within the clade Titanosauria. It includes some of the largest and heaviest dinosaurs known. Description Lognkos ...
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Lithostrotia
Lithostrotia is a clade of derived titanosaur sauropods that lived during the Early Cretaceous and Late Cretaceous. The group was defined by Upchurch ''et al.'' in 2004 as the most recent common ancestor of ''Malawisaurus'' and ''Saltasaurus'' and all the descendants of that ancestor. Lithostrotia is derived from the Ancient Greek , meaning "inlaid with stones", referring to the fact that many known lithostrotians are preserved with osteoderms. However, osteoderms are not a distinguishing feature of the group, as the two noted by Unchurch ''et al.'' include caudal vertebrae with strongly concave front faces (procoely), although the farthest vertebrae are not procoelous. History of research In 1895, Richard Lydekker named the family Titanosauridae to summarize sauropods with procoelous (concave on the front) caudal vertebrae. The name Titanosauridae has since been widely used, and was defined by Salgado and colleagues (1997), Gonzalaz-Riga (2003), and Salgado (2003) as a node-based ...
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Ampelosaurus Atacis
''Ampelosaurus'' ( ; meaning "vine lizard") is a titanosaurian sauropod dinosaur from the Late Cretaceous Period of what is now France. Its type species is ''A. atacis'', named by Le Loeuff in 1995. A possible unnamed species has given ''Ampelosaurus'' an age reaching to the latest Cretaceous, from about 70 to 66 million years ago. Like most sauropods, it would have had a long neck and tail but it also carried armor in the form of osteoderms. Over 500 bones have been assigned to ''Ampelosaurus'' and all but the braincase (assigned to ''A.'' sp.) has been assigned to ''A. atacis''. They are assigned to the same species because all the differences in the limb proportions have been linked to individual variation. ''A. atacis'' is known from a few, well-preserved teeth and some cranial material. A right scapula was discovered associated with a coracoid. The blade of the scapula, contrary to most titanosaurs, is triangular. The blade narrows at one end instead of showing an expans ...
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Ampelosaurus
''Ampelosaurus'' ( ; meaning "vine lizard") is a titanosaurian sauropod dinosaur from the Late Cretaceous Period of what is now France. Its type species is ''A. atacis'', named by Le Loeuff in 1995. A possible unnamed species has given ''Ampelosaurus'' an age reaching to the latest Cretaceous, from about 70 to 66 million years ago. Like most sauropods, it would have had a long neck and tail but it also carried armor in the form of osteoderms. Over 500 bones have been assigned to ''Ampelosaurus'' and all but the braincase (assigned to ''A.'' sp.) has been assigned to ''A. atacis''. They are assigned to the same species because all the differences in the limb proportions have been linked to individual variation. ''A. atacis'' is known from a few, well-preserved teeth and some cranial material. A right scapula was discovered associated with a coracoid. The blade of the scapula, contrary to most titanosaurs, is triangular. The blade narrows at one end instead of showing an expans ...
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Garrigatitan
''Garrigatitan'' (meaning " garrigue giant") is a genus of titanosaurian dinosaur from the late Cretaceous Period of the Grès à Reptiles Formation in France. The genus contains a single species, ''Garrigatitan meridionalis.'' Discovery and naming Between 2009 and 2012, excavations were carried out at Velaux-La Bastide Neuve by the ''Palaios'' Association and the University of Poitiers. During the excavations, the holotype of ''Garrigatitan'' was discovered along with the remains of ''Atsinganosaurus,'' another titanosaurian. In 2021, the type species ''Garrigatitan meridionalis'' was named and described by Verónica Díez Díaz, Géraldine Garcia, Xabier Pereda Suberbiola, Benjamin Jentgen-Ceschino, Koen Stein, Pascal Godefroit and Xavier Valentin. The holotype, MMS / VBN.09.17, was found in a layer of sandstone of the ''Begudian'', the second level of the second series, dating back to the late Campanian. It consists of a sacrum belonging to an immature individual. Additi ...
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Epachthosaurus
''Epachthosaurus'' (meaning "heavy lizard") was a genus of titanosaurian sauropod dinosaur from the Late Cretaceous. It was a basal lithostrotian titanosaur. Its fossils have been found in Central and Northern Patagonia in South America. Discovery and naming The type species, ''E. sciuttoi'', was described by Powell in 1990. The bones assigned to it by Powell in 1990 were, originally, assigned to '' Antarctosaurus sp.'', and then to ''Argyrosaurus superbus?'', before being named as a new taxon. The holotype specimen is MACN-CH 1317, which consists of an incomplete posterior dorsal vertebra. Another specimen, the paratype MACN-CH 18689, consists of a cast of six articulated caudal vertebrae, the partial sacrum, and a fragmentary pubic peduncle from the right ilium. A nearly complete specimen referred to ''Epachthosaurus'', UNPSJB-PV 920, was recovered during field research conducted as part of the project ''Los vertebrados de la Formación Bajo Barreal, Provincia de Chubut, ...
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Atsinganosaurus
''Atsinganosaurus'' is a genus of titanosaurian sauropod dinosaur which existed in what is now France during the late Cretaceous period. Well-preserved remains (and the only known) of ''Atsinganosaurus'' were collected from the Grès à Reptiles Formation of the Aix-en-Provence Basin. ''Atsinganosaurus'' was first described by Géraldine Garcia, Sauveur Amico, Francois Fournier, Eudes Thouand and Xavier Valentin in 2010, and the type and only species is ''Atsinganosaurus velauciensis''. The generic name is derived from the Greek word "τσιγγάνος" or "αθίγγανος", both meaning "gypsy", which refers to the possible migration from east to west of the species. The specific name Specific name may refer to: * in Database management systems, a system-assigned name that is unique within a particular database In taxonomy, either of these two meanings, each with its own set of rules: * Specific name (botany), the two-part (bino ... is named after its finding place, Ve ...
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Lirainosaurus
''Lirainosaurus'' (meaning "slender lizard"; from the Basque ''lirain'', meaning "slender", and the Greek ''sauros'', meaning "lizard")is a genus of titanosaur sauropod which lived in what is now Spain. The type species, ''Lirainosaurus astibiae'', was described by Sanz, Powell, Le Loeuff, Martinez, and Pereda-Suberbiola in 1999. It was a relatively small sauropod, measuring long, possibly up to long for the largest individuals, and weighed about . Classification This genus was based on a skull fragment, isolated teeth, several vertebrae including the holotype - an anterior caudal vertebra, and appendicular bones from the Late Cretaceous of Laño (northern Spain). New material from Laño, Spain described by Diaz ''et al.'' (2013), which includes cervical, dorsal and caudal vertebrae, dorsal ribs, and a haemal arch, has been assigned to ''Lirainosaurus''. According to Diaz ''et al.'', ''Lirainosaurus'' can be distinguished by the presence of a lamina in the interzygapophyseal ...
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Late Campanian
The Campanian is the fifth of six ages of the Late Cretaceous Epoch on the geologic timescale of the International Commission on Stratigraphy (ICS). In chronostratigraphy, it is the fifth of six stages in the Upper Cretaceous Series. Campanian spans the time from 83.6 (± 0.2) to 72.1 (± 0.2) million years ago. It is preceded by the Santonian and it is followed by the Maastrichtian. The Campanian was an age when a worldwide sea level rise covered many coastal areas. The morphology of some of these areas has been preserved: it is an unconformity beneath a cover of marine sedimentary rocks. Etymology The Campanian was introduced in scientific literature by Henri Coquand in 1857. It is named after the French village of Champagne in the department of Charente-Maritime. The original type locality was a series of outcrop near the village of Aubeterre-sur-Dronne in the same region. Definition The base of the Campanian Stage is defined as a place in the stratigraphic column where ...
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Saltasauridae
Saltasauridae (named after the Salta region of Argentina where they were first found) is a family of armored herbivorous sauropods from the Upper Cretaceous. They are known from fossils found in South America, Asia, North America, and Europe. They are characterized by their vertebrae and feet, which are similar to those of ''Saltasaurus'', the first of the group to be discovered and the source of the name. The last and largest of the group and only one found in North America, '' Alamosaurus'', was in length and one of the last sauropods to go extinct. Most of the saltasaurids were smaller, around in length, and one, '' Rocasaurus'', was only long. Like all sauropods, the saltasaurids were quadrupeds, their necks and tails were held almost parallel to the ground, and their small heads had only tiny, peg-like teeth. They were herbivorous, stripping leaves off of plants and digesting them in their enormous guts. Although large animals, they were smaller than other sauropods of th ...
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Aeolosaurini
Aeolosaurini is an extinct clade of titanosaurian dinosaurs known from the Cretaceous period of Argentina and Brazil. Rodrigo M. Santucci and Antonio C. de Arruda-Campos (2011) in their cladistic analysis found ''Aeolosaurus'', ''Gondwanatitan'', '' Maxakalisaurus'', '' Panamericansaurus'' and '' Rinconsaurus'' to be aeolosaurids. Aeolosaurini is characterized by several synapomorphies of the caudal vertebrae, such as angled centra, elongate prezygapophyses, and neural arches shifted anteriorly relative to the centra. In life, their tails may have been strongly curved downward as a result of these traits, which may have increased the force exerted by the caudofemoralis longus muscle in retracting the hindlimb. Some aeolosaurins, such as ''Shingopana'' and ''Overosaurus'', were relatively small compared to other titanosaurs, whereas others, such as ''Aeolosaurus maximus'', were large. Phylogeny Aeolosaurini was defined by Franco-Rosas, Salgado, Rosas and Carvalho (2004) as the ste ...
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Lognkosauria
Lognkosauria is a clade of giant long-necked sauropod dinosaurs within the clade Titanosauria. It includes some of the largest and heaviest dinosaurs known. Description Lognkosaurians can be distinguished from other titanosaurs by the wide and unusually thick cervical rib loops on their neck vertebrae, their extremely robust neck neural spines, the relatively narrow neural canal, and their huge vaulted neural arches. They also had very wide dorsal vertebrae with wing-like side processes, and extremely wide rib cages. Their dorsal side processes are also fairly in-line with the level of the neural canal, instead of being attached further up the neural arch as in lithostrotians. Skull material from ''Malawisaurus'', the sister taxon to Lognkosauria, indicates that lognkosaurians at least began with the big-nosed, rounded head shape of earlier titanosaurs and more basal macronarians. Classification Lognkosauria was defined as the clade encompassing the most recent common ancestor o ...
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Alamosaurus
''Alamosaurus'' (; meaning "Ojo Alamo lizard") is a genus of opisthocoelicaudiine titanosaurian sauropod dinosaurs, containing a single known species, ''Alamosaurus sanjuanensis'', from the late Cretaceous Period of what is now southern North America. Isolated vertebrae and limb bones indicate that it reached sizes comparable to ''Argentinosaurus'' and ''Puertasaurus'', which would make it the largest dinosaur known from North America. Its fossils have been recovered from a variety of rock formations spanning the Maastrichtian age of the late Cretaceous period. Specimens of a juvenile ''Alamosaurus sanjuanensis'' have been recovered from only a few meters below the Cretaceous-Paleogene boundary in Texas, making it among the last surviving non-avian dinosaur species. Description ''Alamosaurus'' was a gigantic quadrupedal herbivore with a long neck and tail and relatively long limbs. Its body was at least partly covered in bony armor. In 2012 Thomas Holtz gave a total length of ...
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