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Limpet
Limpets are a group of aquatic snails that exhibit a conical shell shape (patelliform) and a strong, muscular foot. Limpets are members of the class Gastropoda, but are polyphyletic, meaning the various groups called "limpets" descended independently from different ancestral gastropods. This general category of conical shell is known as "patelliform" (dish-shaped). All members of the large and ancient marine clade Patellogastropoda are limpets. Within that clade, the members of the Patellidae family in particular are often referred to as "true limpets". Other groups, not in the same family, are also called limpets of one type or another, due to the similarity of their shells' shape. Examples include the Fissurellidae ("keyhole limpet") family, which is part of the Vetigastropoda clade (many other members of the Vetigastropoda do not have the morphology of limpets) and the Siphonariidae ("false limpets"), which use a siphon to pump water over their gills. Behaviour and ecolo ...
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Patellogastropoda
The Patellogastropoda, common name true limpet Limpets are a group of aquatic snails that exhibit a conical shell shape (patelliform) and a strong, muscular foot. Limpets are members of the class Gastropoda, but are polyphyletic, meaning the various groups called "limpets" descended indep ...s and historically called the Docoglossa, are members of a major phylogenetic group of marine (ocean), marine gastropods, treated by experts either as a clade or as a taxonomic Order (biology), order. The clade Patellogastropoda is deemed monophyletic based on phylogenetic analysis. Taxonomy Patellogastropoda was proposed by David R. Lindberg, 1986, as an order, and was later included in the subclass Eogastropoda Ponder & Lindberg, 1996. 2005 taxonomy Taxonomy of the Gastropoda (Bouchet & Rocroi, 2005), Bouchet & Rocroi, 2005 designated Patellogastropoda, true limpets, as a clade, rather than as a taxon, but within included superfamilies and families as listed below. Families that a ...
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Fissurellidae
Fissurellidae, common name the keyhole limpets and slit limpets, is a taxonomic family of limpet-like sea snails, marine gastropod molluscs in the clade Vetigastropoda.Rosenberg, G. (2012). Fissurellidae. Accessed through: World Register of Marine Species at http://www.marinespecies.org/aphia.php?p=taxdetails&id=111 on 2013-01-02 Their common name derives from the small hole in the apex of their cone-like shells. Although superficially resembling "true" limpets, they are in fact not closely related to them. Distribution The distribution of fissurellids is worldwide, from cold waters to tropical waters. Powell A. W. B., ''New Zealand Mollusca'', William Collins Publishers Ltd, Auckland, New Zealand 1979 Habitat Fissurellids live in habitats on and under rocks in the lower intertidal zones to deeper waters. Shell description Keyhole limpets somewhat resemble true limpets because of the simple conical shape of their shells, but in reality they are not closely related to true ...
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Common Limpet
''Patella vulgata'', common name the common limpet or common European limpet is a species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ... of sea snail. It is a typical true limpet; a Marine (ocean), marine gastropod mollusc in the family (biology), family Patellidae, with gills.Gofas, S. (2014). Patella vulgata Linnaeus, 1758. Accessed through: World Register of Marine Species at http://www.marinespecies.org/aphia.php?p=taxdetails&id=140685 on 2014-10-29 This species occurs in the waters of Western Europe. Radula The radula in this species is longer than the shell itself. It contains 1,920 teeth in 160 rows of 12 teeth each. ''Patella vulgata'' is found attached to firm substrates from the high shore to the edge of the sublittoral zone, although it predominates in areas of wave ...
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Patellidae
Patellidae is a taxonomic family of sea snails or true limpets, marine gastropod molluscs in the clade Patellogastropoda. (The superfamily Patelloidea should not be confused with the similar-sounding genus of true limpets ''Patelloida'' which is in the family Lottiidae within the superfamily Lottioidea, also part of the Patellogastropoda.) Taxonomy A cladogram showing phylogenic relations of Patellogastropoda based on molecular phylogeny research by Nakano & Ozawa (2007):Nakano T. & Ozawa T. (2007). "Worldwide phylogeography of limpets of the order Patellogastropoda: Molecular, morphological and palaeontological evidence". ''Journal of Molluscan Studies'' 73(1) 79-99. . Genera Genera and species in the family Patellidae include: * † '' Berlieria'' de Loriol, 1903 * ''Cymbula'' H. & A. Adams, 1854 * ''Helcion'' Montfort, 1810 * ''Patella'' Linnaeus, 1758 * † ''Proscutum'' P. Fischer, 1885 * ''Scutellastra'' H. Adams & A. Adams, 1854 ;Synonyms: * ''Ancistromesus'' ...
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Gastropoda
The gastropods (), commonly known as snails and slugs, belong to a large taxonomic class of invertebrates within the phylum Mollusca called Gastropoda (). This class comprises snails and slugs from saltwater, from freshwater, and from land. There are many thousands of species of sea snails and slugs, as well as freshwater snails, freshwater limpets, and land snails and slugs. The class Gastropoda contains a vast total of named species, second only to the insects in overall number. The fossil history of this class goes back to the Late Cambrian. , 721 families of gastropods are known, of which 245 are extinct and appear only in the fossil record, while 476 are currently extant with or without a fossil record. Gastropoda (previously known as univalves and sometimes spelled "Gasteropoda") are a major part of the phylum Mollusca, and are the most highly diversified class in the phylum, with 65,000 to 80,000 living snail and slug species. The anatomy, behavior, feeding, a ...
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Gastropod Shell
The gastropod shell is part of the body of a Gastropoda, gastropod or snail, a kind of mollusc. The shell is an exoskeleton, which protects from predators, mechanical damage, and dehydration, but also serves for muscle attachment and calcium storage. Some gastropods appear shell-less (slugs) but may have a remnant within the mantle, or in some cases the shell is reduced such that the body cannot be retracted within it (semi-slug). Some snails also possess an operculum that seals the opening of the shell, known as the Aperture (mollusc), aperture, which provides further protection. The study of mollusc shells is known as conchology. The biological study of gastropods, and other molluscs in general, is malacology. Shell morphology terms vary by species group. Shell layers The gastropod shell has three major layers secreted by the Mantle (mollusc), mantle. The calcareous central layer, tracum, is typically made of calcium carbonate precipitated into an organic matrix known as c ...
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Siphonariidae
Siphonariidae, also known as false limpets, are a taxonomic family of small to medium-sized air-breathing sea snails, marine and brackish water pulmonate gastropod molluscs. __TOC__ Shell description Empty siphonariid limpet shells can be distinguished from true limpet shells by examining the interior. In the siphonariids there is a well-marked lateral groove on the right side of the shell, and a corresponding interruption of the ring of muscle attachment scars. These markers show the positioning of the pneumostome or pulmonary The lungs are the primary organs of the respiratory system in humans and most other animals, including some snails and a small number of fish. In mammals and most other vertebrates, two lungs are located near the backbone on either side of ... orifice. Genera Genera in the family Siphonariidae include: * '' Anthosiphonaria'' Kuroda & Habe, 1971 ** '' Anthosiphonaria sirius'' * '' Aporemodon'' Robson, 1913 * ''Benhamina'' Finlay, 1927 - wi ...
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Radula
The radula (, ; plural radulae or radulas) is an anatomical structure used by molluscs for feeding, sometimes compared to a tongue. It is a minutely toothed, chitinous ribbon, which is typically used for scraping or cutting food before the food enters the esophagus. The radula is unique to the molluscs, and is found in every class of mollusc except the bivalves, which instead use cilia, waving filaments that bring minute organisms to the mouth. Within the gastropods, the radula is used in feeding by both herbivorous and carnivorous snails and slugs. The arrangement of teeth ( denticles) on the radular ribbon varies considerably from one group to another. In most of the more ancient lineages of gastropods, the radula is used to graze, by scraping diatoms and other microscopic algae off rock surfaces and other substrates. Predatory marine snails such as the Naticidae use the radula plus an acidic secretion to bore through the shell of other molluscs. Other predatory marine snails ...
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Vetigastropoda
Vetigastropoda is a major Taxonomy (biology), taxonomic group of sea snails, marine (ocean), marine gastropod mollusc, mollusks that form a very ancient lineage (evolution), lineage. Taxonomically the Vetigastropoda are sometimes treated as an Order (biology), order, although they are treated as an unranked clade in taxonomy of the Gastropoda (Bouchet & Rocroi, 2005), Bouchet and Rocroi, 2005. Vetigastropods are considered to be among the most primitive living gastropods, and are widely distributed in all oceans of the world. Their habitats range from the deep sea to intertidal zones. Many have exoskeleton, shells with slits or other secondary openings. One of their main characteristics is the presence of intersected crossed platy shell structure. Most vetigastropods have some bilateral asymmetry of their Organ (anatomy), organ systems. Description Vetigastropods range in size from approximately 0.08 in (2 mm) long in the case of Scissurelloidea or Skeneoidea, to more than ...
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Ventricle (heart)
A ventricle is one of two large chambers toward the bottom of the heart that collect and expel blood towards the peripheral beds within the body and lungs. The blood pumped by a ventricle is supplied by an atrium, an adjacent chamber in the upper heart that is smaller than a ventricle. Interventricular means between the ventricles (for example the interventricular septum), while intraventricular means within one ventricle (for example an intraventricular block). In a four-chambered heart, such as that in humans, there are two ventricles that operate in a double circulatory system: the right ventricle pumps blood into the pulmonary circulation to the lungs, and the left ventricle pumps blood into the systemic circulation through the aorta. Structure Ventricles have thicker walls than atria and generate higher blood pressures. The physiological load on the ventricles requiring pumping of blood throughout the body and lungs is much greater than the pressure generated by the atria ...
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Circulatory System
The blood circulatory system is a system of organs that includes the heart, blood vessels, and blood which is circulated throughout the entire body of a human or other vertebrate. It includes the cardiovascular system, or vascular system, that consists of the heart and blood vessels (from Greek ''kardia'' meaning ''heart'', and from Latin ''vascula'' meaning ''vessels''). The circulatory system has two divisions, a systemic circulation or circuit, and a pulmonary circulation or circuit. Some sources use the terms ''cardiovascular system'' and ''vascular system'' interchangeably with the ''circulatory system''. The network of blood vessels are the great vessels of the heart including large elastic arteries, and large veins; other arteries, smaller arterioles, capillaries that join with venules (small veins), and other veins. The Closed circulatory system, circulatory system is closed in vertebrates, which means that the blood never leaves the network of blood vessels. Some in ...
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Heart
The heart is a muscular organ in most animals. This organ pumps blood through the blood vessels of the circulatory system. The pumped blood carries oxygen and nutrients to the body, while carrying metabolic waste such as carbon dioxide to the lungs. In humans, the heart is approximately the size of a closed fist and is located between the lungs, in the middle compartment of the chest. In humans, other mammals, and birds, the heart is divided into four chambers: upper left and right atria and lower left and right ventricles. Commonly the right atrium and ventricle are referred together as the right heart and their left counterparts as the left heart. Fish, in contrast, have two chambers, an atrium and a ventricle, while most reptiles have three chambers. In a healthy heart blood flows one way through the heart due to heart valves, which prevent backflow. The heart is enclosed in a protective sac, the pericardium, which also contains a small amount of fluid. The wall ...
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