Loxodidae
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Loxodidae
Loxodidae is a family of karyorelict ciliates. Loxodidae members possess an elongated, laterally flattened shape. They share two key characters: a beak-like anterior rostrum interrupting the perioral kineties, and peculiar cytoplasmic organelles named Müller vesicles. The extensive development of lacunae of the smooth endoplasmic reticulum leads to strong vacuolization of the endoplasm. This feature is associated to a lack of contractile vacuoles in all loxodids. The term ''Loxodidae'' derives from the ancient Greek (), meaning "oblique, tilted". Gravitaxis Loxodidae members possess the ability to orient themselves in oxygen gradients. They use gravity as a stimulus for this spatial orientation, a phenomenon called gravitaxis or geotaxis. Loxodid ciliates must therefore have developed mechanoreceptors informing them about what is up or down. A likely candidate structure for their gravitaxis is the Müller vesicle. Müller vesicle Müller vesicles (also known as Müller ...
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Remanella
''Remanella'' is a genus of karyorelict ciliates, belonging to family Loxodidae. Whereas ''Remanella'' inhabits brackish and marine waters, ''Loxodes'' – the other loxodid genus – is a freshwater taxon. Etymology The genus name is a patronym honoring the German zoologist Adolf Remane (1898–1976), who contributed to the discovery of the marine interstitial fauna. Phylogeny Molecular phylogeny based on sequences of the SSU rRNA gene suggests that the genus ''Remanella'' might be paraphyletic with respect to a monophyletic In cladistics for a group of organisms, monophyly is the condition of being a clade—that is, a group of taxa composed only of a common ancestor (or more precisely an ancestral population) and all of its lineal descendants. Monophyletic gro ... genus ''Loxodes'' but this result is not strongly supported. Species list At least 18 species are recognized in the genus ''Remanella'': References External links * ' {{Taxonbar, from=Q25 ...
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Loxodes
''Loxodes'' is a genus of karyorelictean ciliates, belonging to family Loxodidae. It is the only known karyorelictean ciliate that lives in freshwater habitats. The term ''Loxodes'' derives from the ancient greek (), meaning "oblique, tilted". Ecology ''Loxodes'' lives in freshwater habitats such as lakes and ponds, unlike other karyorelictean ciliates such as the other loxodid genus ''Remanella'', which live in brackish-water or marine habitats. They feed on bacteria and protists such as microalgae. It is microaerobic, preferring low concentrations of oxygen, below 5% atmospheric saturation. It can also survive extended periods in anoxic water, where oxygen is absent. Under such conditions, ''Loxodes'' is able to use nitrate instead of oxygen as an electron acceptor for respiration. Nitrate respiration is rare among eukaryotes, and ''Loxodes'' was the first eukaryote known to have this capability. ''Loxodes'' is also sensitive to light. Geotaxis Both genera in the fam ...
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Statocyst
The statocyst is a balance sensory receptor present in some aquatic invertebrates, including bivalves, cnidarians, ctenophorans, echinoderms, cephalopods, and crustaceans. A similar structure is also found in ''Xenoturbella''. The statocyst consists of a sac-like structure containing a mineralised mass (statolith) and numerous innervated sensory hairs (setae). The statolith's inertia causes it to push against the setae when the animal accelerates. Deflection of setae by the statolith in response to gravity activates neurons, providing feedback to the animal on change in orientation and allowing balance to be maintained. In other words, the statolith shifts as the animal moves. Any movement large enough to throw the organism off balance causes the statolith to brush against tiny bristles which in turn send a message to the brain to correct its balance. It may have been present in the common ancestor of cnidarians and bilaterians. Hearing In cephalopods like squids, statocysts ...
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Karyorelictea
Karyorelictea is a class of ciliates in the subphylum Postciliodesmatophora. Most species are members of the microbenthos community, that is, microscopic organisms found in the marine interstitial habitat, though one genus, ''Loxodes'', is found in freshwater. The majority of karyorelict taxa have not been cultivated in the laboratory, although clonal lines of ''Loxodes'' have been developed. Systematics According to Lynn (2008), the Karyorelictea class is divided into three orders: * Loxodida, containing the families Cryptopharyngidae and Loxodidae; * Protoheterotrichida, containing the families Aveliidae and Geleiidae; * Protostomatida, containing the families Kentrophoridae and Trachelocercidae. These three orders were defined morphologically, and have been confirmed with molecular phylogenetics. An additional family, Wilbertomorphidae, is of uncertain affiliation and has not been assigned to an order. Nuclear dimorphism All ciliates, including karyorelicteans, p ...
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Ancient Greek
Ancient Greek includes the forms of the Greek language used in ancient Greece and the ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Dark Ages (), the Archaic period (), and the Classical period (). Ancient Greek was the language of Homer and of fifth-century Athenian historians, playwrights, and philosophers. It has contributed many words to English vocabulary and has been a standard subject of study in educational institutions of the Western world since the Renaissance. This article primarily contains information about the Epic and Classical periods of the language. From the Hellenistic period (), Ancient Greek was followed by Koine Greek, which is regarded as a separate historical stage, although its earliest form closely resembles Attic Greek and its latest form approaches Medieval Greek. There were several regional dialects of Ancient Greek, of which Attic Greek developed into Koine. Dia ...
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Drawing Of Loxodes Striatus By E Penard
Drawing is a visual art that uses an instrument to mark paper or another two-dimensional surface. The instruments used to make a drawing are pencils, crayons, pens with inks, brushes with paints, or combinations of these, and in more modern times, computer styluses with graphics tablets or gamepads in VR drawing software. A drawing instrument releases a small amount of material onto a surface, leaving a visible mark. The most common support for drawing is paper, although other materials, such as cardboard, vellum, wood, plastic, leather, canvas, and board, have been used. Temporary drawings may be made on a blackboard or whiteboard. Drawing has been a popular and fundamental means of public expression throughout human history. It is one of the simplest and most efficient means of communicating ideas. The wide availability of drawing instruments makes drawing one of the most common artistic activities. In addition to its more artistic forms, drawing is frequently used in commer ...
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Gravity
In physics, gravity () is a fundamental interaction which causes mutual attraction between all things with mass or energy. Gravity is, by far, the weakest of the four fundamental interactions, approximately 1038 times weaker than the strong interaction, 1036 times weaker than the electromagnetic force and 1029 times weaker than the weak interaction. As a result, it has no significant influence at the level of subatomic particles. However, gravity is the most significant interaction between objects at the macroscopic scale, and it determines the motion of planets, stars, galaxies, and even light. On Earth, gravity gives weight to physical objects, and the Moon's gravity is responsible for sublunar tides in the oceans (the corresponding antipodal tide is caused by the inertia of the Earth and Moon orbiting one another). Gravity also has many important biological functions, helping to guide the growth of plants through the process of gravitropism and influencing the circ ...
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Stimulus (physiology)
In physiology, a stimulus is a detectable change in the physical or chemical structure of an organism's internal or external environment. The ability of an organism or organ to detect external stimuli, so that an appropriate reaction can be made, is called sensitivity (excitability). Sensory receptors can receive information from outside the body, as in touch receptors found in the skin or light receptors in the eye, as well as from inside the body, as in chemoreceptors and mechanoreceptors. When a stimulus is detected by a sensory receptor, it can elicit a reflex via stimulus transduction. An internal stimulus is often the first component of a homeostatic control system. External stimuli are capable of producing systemic responses throughout the body, as in the fight-or-flight response. In order for a stimulus to be detected with high probability, its level of strength must exceed the absolute threshold; if a signal does reach threshold, the information is transmitted to ...
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Gravitaxis
Gravitaxis (or ''geotaxis'') is a form of taxis characterized by the directional movement of an organism in response to gravity. Gravitaxis is one of the many forms of taxis. It is characterized by the movement of an organism in response to gravitational forces. It is sometimes called geotaxis. There are a few different causes for gravitaxis. Many microorganisms have receptors like statocysts that allow them to sense the direction of gravity and to adjust their orientation accordingly. However, gravitaxis can result also from a purely physical mechanism so that organs for sensing the direction of gravity are not necessary. An example is given by microorganisms with a center of mass that is shifted to one end of the organism. Similar to a buoy, such mass-anisotropic microorganisms orient upwards under gravity. It has been shown that even an asymmetry in the shape of microorganisms can be sufficient to cause gravitaxis. Gravitaxis is different from gravitropism in a way that the latt ...
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Geotaxis
Gravitaxis (or ''geotaxis'') is a form of taxis characterized by the directional movement of an organism in response to gravity. Gravitaxis is one of the many forms of taxis. It is characterized by the movement of an organism in response to gravitational forces. It is sometimes called geotaxis. There are a few different causes for gravitaxis. Many microorganisms have receptors like statocysts that allow them to sense the direction of gravity and to adjust their orientation accordingly. However, gravitaxis can result also from a purely physical mechanism so that organs for sensing the direction of gravity are not necessary. An example is given by microorganisms with a center of mass that is shifted to one end of the organism. Similar to a buoy, such mass-anisotropic microorganisms orient upwards under gravity. It has been shown that even an asymmetry in the shape of microorganisms can be sufficient to cause gravitaxis. Gravitaxis is different from gravitropism in a way that the latt ...
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Smooth Endoplasmic Reticulum
The endoplasmic reticulum (ER) is, in essence, the transportation system of the eukaryotic cell, and has many other important functions such as protein folding. It is a type of organelle made up of two subunits – rough endoplasmic reticulum (RER), and smooth endoplasmic reticulum (SER). The endoplasmic reticulum is found in most eukaryotic cells and forms an interconnected network of flattened, membrane-enclosed sacs known as cisternae (in the RER), and tubular structures in the SER. The membranes of the ER are continuous with the outer nuclear membrane. The endoplasmic reticulum is not found in red blood cells, or spermatozoa. The two types of ER share many of the same proteins and engage in certain common activities such as the synthesis of certain lipids and cholesterol. Different types of cells contain different ratios of the two types of ER depending on the activities of the cell. RER is found mainly toward the nucleus of cell and SER towards the cell membrane or plasma ...
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Otto Friedrich Müller
Otto Friedrich Müller, also known as Otto Friedrich Mueller (2 November 1730 – 26 December 1784) was a Danish naturalist and scientific illustrator. Biography Müller was born in Copenhagen. He was educated for the church, became tutor to a young nobleman, and after several years' travel with him, settled in Copenhagen in 1767, and married a lady of wealth. His first important works, ''Fauna Insectorum Friedrichsdaliana'' (Leipzig, 1764), and ''Flora Friedrichsdaliana'' (Strasbourg, 1767), giving accounts of the insects and flora of the estate of Frederiksdal, near Copenhagen, recommended him to Frederick V of Denmark, by whom he was employed to continue the ''Flora Danica'' a comprehensive atlas of the flora of Denmark. Müller added two volumes to the three published by Georg Christian Oeder since 1761. The study of invertebrates began to occupy his attention almost exclusively, and in 1771 he produced a work in German on “Certain Worms inhabiting Fresh and Salt Water,†...
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