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Lancian
The Lancian was a North American faunal stage of the Late Cretaceous. It was the final stage of the Cretaceous period in North America, lasting from approximately 70.6 to 66 million years ago. Geology Terrestrial sedimentary strata from the Judithian to the Lancian are generally regressive throughout the entire sequence, so the preserved changes in fossil communities represent not only phylogenetic changes but ecological zones from the submontane habitats to near-sea level coastal habitats. Paleobiogeography By the Lancian, the crested hadrosaurs are no longer the dominant inhabitant of any province of western North America; the only remaining species was '' Hypacrosaurus''. Lehman records three surviving chasmosaurs, ''Triceratops,'' '' Torosaurus'' and ''Nedoceratops'', with the possibility of the recently discovered ''Ojoceratops, Regaliceratops and'' ''Bravoceratops''. It has recently been suggested that ''Triceratops'' and ''Torosaurus'' may be synonymous, though this is sti ...
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Triceratops
''Triceratops'' ( ; ) is a genus of herbivore, herbivorous Chasmosaurinae, chasmosaurine Ceratopsidae, ceratopsid dinosaur that first appeared during the late Maastrichtian stage of the Late Cretaceous Period (geology), period, about 68 million years ago in what is now North America. It is one of the last-known non-avian dinosaur genera, and became extinct in the Cretaceous-Paleogene extinction event, Cretaceous–Paleogene extinction event 66 million years ago. The name ''Triceratops'', which literally means 'three-horned face', is derived from the Ancient Greek, Greek words () meaning 'three', () meaning 'horn', and () meaning 'face'. Bearing a large bony neck frill, frill, three horn (anatomy), horns on the skull, and a large four-legged body, exhibiting convergent evolution with rhinoceroses and bovines, ''Triceratops'' is one of the most recognizable of all dinosaurs and the most well-known ceratopsid. It was also one of the largest, up to long and in body m ...
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Edmontosaurus Annectens
''Edmontosaurus annectens'' (meaning "connected lizard from Edmonton") is a species of flat-headed and duck-billed (hadrosaurid) dinosaur from the very end of the Cretaceous Period (geology), Period, in what is now North America. Remains of ''E. annectens'' have been preserved in the Frenchman Formation, Frenchman, Hell Creek Formation, Hell Creek, and Lance Formations. All of these formations are dated to the late Maastrichtian stage of the Late Cretaceous Period, representing the last three million years before the Cretaceous–Paleogene boundary, extinction of the dinosaurs (between 68 and 66 million years agoHoltz, Thomas R. Jr. (2012) ''Dinosaurs: The Most Complete, Up-to-Date Encyclopedia for Dinosaur Lovers of All Ages,'Winter 2011 Appendix./ref>). ''E. annectens'' is also found in the Laramie Formation, and magnetostratigraphy suggests an age of 69-68 Ma for the Laramie Formation.*Hicks, J.F., Johnson, K.R., Obradovich, J. D., Miggins, D.P., and Tauxe, L. 2003. Magnet ...
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Quetzalcoatlus
''Quetzalcoatlus'' is a genus of pterosaur known from the Late Cretaceous period of North America (Maastrichtian stage); its members were among the largest known flying animals of all time. ''Quetzalcoatlus'' is a member of the Azhdarchidae, a family of advanced toothless pterosaurs with unusually long, stiffened necks. Its name comes from the Aztec feathered serpent god Quetzalcoatl. The type species is ''Q. northropi'', named by Douglas Lawson in 1975; the genus also includes the smaller species ''Q. lawsoni'', which was known for many years as an unnamed species before being named by Brian Andres and Wann Langston Jr. (posthumously) in 2021. Discovery and species The first ''Quetzalcoatlus'' fossils were discovered in Texas, United States, from the Maastrichtian Javelina Formation at Big Bend National Park (dated to around 68 million years ago) in 1971 by Douglas A. Lawson, then a geology graduate student from the Jackson School of Geosciences at the University of Texas ...
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Alamosaurus
''Alamosaurus'' (; meaning "Ojo Alamo lizard") is a genus of opisthocoelicaudiine titanosaurian sauropod dinosaurs, containing a single known species, ''Alamosaurus sanjuanensis'', from the late Cretaceous Period of what is now southern North America. Isolated vertebrae and limb bones indicate that it reached sizes comparable to ''Argentinosaurus'' and ''Puertasaurus'', which would make it the largest dinosaur known from North America. Its fossils have been recovered from a variety of rock formations spanning the Maastrichtian age of the late Cretaceous period. Specimens of a juvenile ''Alamosaurus sanjuanensis'' have been recovered from only a few meters below the Cretaceous-Paleogene boundary in Texas, making it among the last surviving non-avian dinosaur species. Description ''Alamosaurus'' was a gigantic quadrupedal herbivore with a long neck and tail and relatively long limbs. Its body was at least partly covered in bony armor. In 2012 Thomas Holtz gave a total length of ...
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Faunal Stage
In chronostratigraphy, a stage is a succession of rock strata laid down in a single age on the geologic timescale, which usually represents millions of years of deposition. A given stage of rock and the corresponding age of time will by convention have the same name, and the same boundaries. Rock series are divided into stages, just as geological epochs are divided into ages. Stages can be divided into smaller stratigraphic units called chronozones. (See chart at right for full terminology hierarchy.) Stages may also be divided into substages or indeed grouped as superstages. The term faunal stage is sometimes used, referring to the fact that the same fauna (animals) are found throughout the layer (by definition). Definition Stages are primarily defined by a consistent set of fossils (biostratigraphy) or a consistent magnetic polarity (see paleomagnetism) in the rock. Usually one or more index fossils that are common, found worldwide, easily recognized, and limited to a sing ...
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Denversaurus
''Denversaurus'' (meaning "Denver lizard") is a genus of panoplosaurin nodosaurid dinosaur from the Late Cretaceous (late Maastrichtian) of western North America. Although at one point treated as a junior synonym of ''Edmontonia'' by some taxonomists, current research indicates that it is a distinct nodosaurid genus. Discovery and naming In 1922, Philip Reinheimer, a collector and technician employed by the Colorado Museum of Natural History, the predecessor of the present Denver Museum of Nature and Science, near the Twito Ranch in Corson County, South Dakota discovered the fossil of an ankylosaurian in a Maastrichtian age terrestrial horizon of the Lance Formation. In 1943, American paleontologist Barnum Brown referred the find to ''Edmontonia longiceps''. In 1988, Robert Thomas Bakker decided to split the genus ''Edmontonia''. The species ''Edmontonia rugosidens'' he made into a separate genus ''Chassternbergia'' and the Denver fossil was named and described as a new genu ...
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Early Cretaceous
The Early Cretaceous ( geochronological name) or the Lower Cretaceous (chronostratigraphic name), is the earlier or lower of the two major divisions of the Cretaceous. It is usually considered to stretch from 145  Ma to 100.5 Ma. Geology Proposals for the exact age of the Barremian-Aptian boundary ranged from 126 to 117 Ma until recently (as of 2019), but based on drillholes in Svalbard the defining early Aptian Oceanic Anoxic Event 1a (OAE1a) was carbon isotope dated to 123.1±0.3 Ma, limiting the possible range for the boundary to c. 122–121 Ma. There is a possible link between this anoxic event and a series of Early Cretaceous large igneous provinces (LIP). The Ontong Java-Manihiki-Hikurangi large igneous province, emplaced in the South Pacific at c. 120 Ma, is by far the largest LIP in Earth's history. The Ontong Java Plateau today covers an area of 1,860,000 km2. In the Indian Ocean another LIP began to form at c. 120 Ma, the Kerguelen P ...
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Avisaurus
''Avisaurus'' (meaning "bird lizard") is a genus of enantiornithine bird from the Late Cretaceous of North America. Discovery ''Avisaurus archibaldi'' was discovered in the Late Cretaceous Hell Creek Formation of North America (Maastrichtian, from c.70.6-66 million years ago), making it one of the last enantiornithids. It was collected in 1975 in the UCMP locality V73097, in Garfield County, Montana, USA. The holotype is represented by a single fossil of a tarsometatarsus in the collection of the University of California Museum of Paleontology. It has the catalog number UCMP 117600. The species name honors J. David Archibald, its discoverer, from The University of California, Berkeley. It was initially described as the left tarsometatarsus of a non-avian theropod by Brett-Surman and Paul in 1985. It was later redescribed as the right tarsometatarsus of an enantiornithine bird by Chiappe in 1992.Chiappe, Luis M. (1992) "Enantiornithine (Aves) Tarsometatarsi and the Avian Affinities ...
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Nodocephalosaurus
''Nodocephalosaurus'' (meaning "knob headed lizard") is a monospecific genus of ankylosaurid dinosaur from New Mexico that lived during the Late Cretaceous (late Campanian to early Maastrichtian stage, 73.49 to 73.04 Ma) in what is now the De-na-zin member of the Kirtland Formation. The type and only species, ''Nodocephalosaurus kirtlandensis'', is known only from a partial skull. It was named in 1999 by Robert M. Sullivan. ''Nodocephalosaurus'' has an estimated length of 4.5 metres (15 feet) and weight of 1.5 tonnes (3,306 lbs).Paul, G.S., 2016, ''The Princeton Field Guide to Dinosaurs 2nd Edition'', Princeton University Press It is closely related and shares similar cranial anatomy to ''Akainacephalus''. Discovery and naming In 1995, a partial skull of an ankylosaur was discovered weathering out of a grey mudstone a few hundred metres west of a new ''Parasaurolophus'' site in the De-na-zin member of the Kirtland Formation, New Mexico. Robert M. Sullivan and Thomas E. William ...
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Neoceratopsian
Ceratopsia or Ceratopia ( or ; Greek: "horned faces") is a group of herbivorous, beaked dinosaurs that thrived in what are now North America, Europe, and Asia, during the Cretaceous Period, although ancestral forms lived earlier, in the Jurassic. The earliest known ceratopsian, ''Yinlong downsi'', lived between 161.2 and 155.7 million years ago.Holtz, Thomas R. Jr. (2011) ''Dinosaurs: The Most Complete, Up-to-Date Encyclopedia for Dinosaur Lovers of All Ages,'Winter 2010 Appendix./ref> The last ceratopsian species, ''Triceratops prorsus'', became extinct during the Cretaceous–Paleogene extinction event, . ''Triceratops'' is by far the best-known ceratopsian to the general public. It is traditional for ceratopsian genus names to end in "''-ceratops''", although this is not always the case. One of the first named genera was ''Ceratops'' itself, which lent its name to the group, although it is considered a ''nomen dubium'' today as its fossil remains have no distinguishing characte ...
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Anagenesis
Anagenesis is the gradual evolution of a species that continues to exist as an interbreeding population. This contrasts with cladogenesis, which occurs when there is branching or splitting, leading to two or more lineages and resulting in separate species. Anagenesis does not always lead to the formation of a new species from an ancestral species. When speciation does occur as different lineages branch off and cease to interbreed, a core group may continue to be defined as the original species. The evolution of this group, without extinction or species selection, is anagenesis. Hypotheses One hypothesis is that during the speciation event in anagenetic evolution, the original populations will increase quickly, and then rack up genetic variation over long periods of time by mutation and recombination in a stable environment. Other factors such as selection or genetic drift will have such a significant effect on genetic material and physical traits that a species can be acknow ...
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Bearpaw Transgression
Bearpaw or Bear Paw may refer to: *Bearpaw (brand), a brand of footwear, including sheepskin boots, slippers and casual shoes *Bearpaw Formation, a rock formation in North America * Bear-paw poppies, the genus of the poppy family Papaveraceae * Mikołaj "Bearpaw" Potocki (1595–1651), Polish nobleman * Bearpaw Mountain, a summit in Washington state *Bear Paw Mountains, mountain range in the U.S. state of Montana *Bear's Paw, a mountain in the U.S. state of North Carolina *a type of snowshoe See also *Battle of Bear Paw The Battle of Bear Paw (also sometimes called Battle of the Bears Paw or Battle of the Bears Paw Mountains) was the final engagement of the Nez Perce War of 1877. Following a running fight from north central Idaho Territory over the previous f ...
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