''Avisaurus'' (meaning "bird lizard") is a

genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...

enantiornithine The Enantiornithes, also known as enantiornithines or enantiornitheans in literature, are a group of extinct avialans ("birds" in the broad sense), the most abundant and diverse group known from the Mesozoic era. Almost all retained teeth and cla ...

bird Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweigh ...

from the

Late Cretaceous The Late Cretaceous (100.5–66 Ma) is the younger of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''creta'', the ...

North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and the Car ...

Discovery

''Avisaurus archibaldi'' was discovered in the Late Cretaceous

Hell Creek Formation The Hell Creek Formation is an intensively studied division of mostly Upper Cretaceous and some lower Paleocene rocks in North America, named for exposures studied along Hell Creek, near Jordan, Montana. The formation stretches over portions of ...

of North America (

Maastrichtian The Maastrichtian () is, in the ICS geologic timescale, the latest age (uppermost stage) of the Late Cretaceous Epoch or Upper Cretaceous Series, the Cretaceous Period or System, and of the Mesozoic Era or Erathem. It spanned the interval from ...

, from c.70.6-66

million years ago The abbreviation Myr, "million years", is a unit of a quantity of (i.e. ) years, or 31.556926 teraseconds. Usage Myr (million years) is in common use in fields such as Earth science and cosmology. Myr is also used with Mya (million years ago). ...

), making it one of the last enantiornithids. It was collected in 1975 in the UCMP locality V73097, in Garfield County, Montana, USA. The holotype is represented by a single fossil of a

tarsometatarsus The tarsometatarsus is a bone that is only found in the lower leg of birds and some non-avian dinosaurs. It is formed from the fusion of several bones found in other types of animals, and homologous to the mammalian tarsus (ankle bones) and meta ...

in the collection of the University of California Museum of Paleontology. It has the catalog number UCMP 117600. The species name honors J. David Archibald, its discoverer, from

The University of California, Berkeley The University of California, Berkeley (UC Berkeley, Berkeley, Cal, or California) is a public land-grant research university in Berkeley, California. Established in 1868 as the University of California, it is the state's first land-grant univ ...

. It was initially described as the left tarsometatarsus of a non-avian

theropod Theropoda (; ), whose members are known as theropods, is a dinosaur clade that is characterized by hollow bones and three toes and claws on each limb. Theropods are generally classed as a group of saurischian dinosaurs. They were ancestrally c ...

by Brett-Surman and Paul in 1985. It was later redescribed as the right tarsometatarsus of an

bird by Chiappe in 1992.Chiappe, Luis M. (1992) "Enantiornithine (Aves) Tarsometatarsi and the Avian Affinities of the Late Cretaceous Avisauridae" "Journal of Vertebrate Paleontology" September 3, 1992, Volume 12 no. 3 pp. 344-350 ''Avisaurus gloriae'' Varricchio and Chiappe 1995, discovered in the late

Campanian The Campanian is the fifth of six ages of the Late Cretaceous Epoch on the geologic timescale of the International Commission on Stratigraphy (ICS). In chronostratigraphy, it is the fifth of six stages in the Upper Cretaceous Series. Campanian s ...

Upper

Two Medicine Formation The Two Medicine Formation is a geological formation, or rock body, in northwestern Montana and southern Alberta that was deposited between and (million years ago), during Campanian (Late Cretaceous) time. It crops out to the east of the Rocky M ...

of Glacier County, Montana, USA, was renamed ''

Gettyia ''Gettyia'' is an extinct genus of avisaurid enantiornithean bird from the Late Cretaceous of North America. Description ''Gettyia'' is known from a single right tarsometatarsus. This bone was similar to that of other North American avisau ...

'' by Atterholt et al. (2018).

Description

The specimen has a maximum length of , making it one of the largest known tarsometatarsi of an enantiornithine.Varricchio, David J., Chiappe, Luis M. (1995) "A New Enantiornithine Bird From the Upper Cretaceous Two medicine Formation of Montana" " Journal of Vertebrate Paleontology" March 14, 1995, Vol. 15, no. 1, pp. 201 - 204 RAM 14306, an incomplete

coracoid A coracoid (from Greek κόραξ, ''koraks'', raven) is a paired bone which is part of the shoulder assembly in all vertebrates except therian mammals (marsupials and placentals). In therian mammals (including humans), a coracoid process is prese ...

, is bigger than the holotype. It indicates an animal with a length of , hip height of , and weight of .

Classification

This genus belongs to the

family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...

Avisauridae, which also contains similar animals from

South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the southe ...

such as ''

Soroavisaurus ''Soroavisaurus'' is a genus of enantiornithean birds related to '' Avisaurus''. It lived during the Late Cretaceous of Argentina. The only known species, ''S. australis'', is known from fossils collected from the Lecho Formation (Maastrichtian ...

'' Chiappe, Luis M. (1993) " Enantiornithine (Aves) Tarsometatarsi from the Cretaceous Lecho Formation of Northwestern Argentina" ''American Museum Novitates'' December 27, 1993 Number 3083 pp. 1-2 and ''

Neuquenornis ''Neuquenornis volans''Chiappe, Luis M.,Calvo, Jorge O. (1994) "Neuquenornis volans, a New Late Cretaceous Bird (Enantiornithes: Avisauridae) from Patagonia, Argentina""Journal of Vertebrate Paleontology" Vol. 14, No. 2 (Jun. 22, 1994), pp. 230- ...

'' Chiappe, Luis M., Calvo, Jorge O. (1994) "Neuquenornis volans, a New Late Cretaceous Bird (Enantiornithes: Avisauridae) from Patagonia, Argentina" "Journal of Vertebrate Paleontology" June 22, 1994 Volume 14 No. 2 pp.230-246. In the Late Cretaceous the

Americas The Americas, which are sometimes collectively called America, are a landmass comprising the totality of North and South America. The Americas make up most of the land in Earth's Western Hemisphere and comprise the New World. Along with th ...

were still separated by a branch of the

Tethys Ocean The Tethys Ocean ( el, Τηθύς ''Tēthús''), also called the Tethys Sea or the Neo-Tethys, was a prehistoric ocean that covered most of the Earth during much of the Mesozoic Era and early Cenozoic Era, located between the ancient continents ...

. The paper by Brett-Surman and Paul in 1985 explicitly considered the possibility that ''A. archibaldi'' was an enantiornithine. The authors described and named UCMP 117600 formally, but they looked at other enantiornithine material, including the "

metatarsus The metatarsal bones, or metatarsus, are a group of five long bones in the foot, located between the tarsal bones of the hind- and mid-foot and the phalanges of the toes. Lacking individual names, the metatarsal bones are numbered from the med ...

" PVL 4690 from Argentina. The authors assigned this latter fossil to ''Avisaurus'' sp. From this they concluded that members of the genus ''Avisaurus'' existed in both North and South America in the Late Cretaceous. Moreover, the authors concluded that the length/width ratio and degree of metacarpal fusion of these bones were more like those of non — avian dinosaurs. A terrestrial dinosaur genus in both continents would then support Brett — Surman's theory that there had been a land connection between the two continents.Brett-Surman, Michael K., Paul, Gregory S. (1985) "A new family of bird-like dinosaurs linking Laurasia and Gondwanaland." "Journal of Vertebrate Paleontology" 5(2): 133-138. Further discoveries and further study by Chiappe showed that all of the material belonged to enantiornithine birds, and PVL 4690 was given its own genus ''

''.

Paleoenvironment

Avisaurus and Brachychampsa by tomozsaurus

''Avisaurus'' is known from the

humid Humidity is the concentration of water vapor present in the air. Water vapor, the gaseous state of water, is generally invisible to the human eye. Humidity indicates the likelihood for precipitation, dew, or fog to be present. Humidity depen ...

low-lying swamps, lakes, and river basins of the western shore of the

Western Interior Seaway The Western Interior Seaway (also called the Cretaceous Seaway, the Niobraran Sea, the North American Inland Sea, and the Western Interior Sea) was a large inland sea that split the continent of North America into two landmasses. The ancient sea, ...

, and from the much more arid uplands between that area and the Cordilleran Overthrust Belt which eventually formed the

Rocky Mountains The Rocky Mountains, also known as the Rockies, are a major mountain range and the largest mountain system in North America. The Rocky Mountains stretch in straight-line distance from the northernmost part of western Canada, to New Mexico in ...

. In those flooded coastal wooded plains with predominantly coniferous trees and flowering bush ''Avisaurus'' were hunting. A strong tarsus of ''Avisaurus'' with inwardly curved claws resembles that of eudromaeosaurs. With these claws, they could grab and hold prey, presumably having a resemblance to modern raptors.

Discovery

Description

Classification

Paleoenvironment

References

Further reading