Kourisodon
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Kourisodon
''Kourisodon'' ("razor tooth") is an extinct genus of mosasaur. Fossils have been found from Vancouver Island in British Columbia, Canada,Nicholls, E. L. and Meckert, D. (2002). Marine reptiles from the Nanaimo Group (Upper Cretaceous) of Vancouver Island. ''Canadian Journal of Earth Science'' 39(11):1591-1603. as well as from the Izumi Group of Japan. These finds date back to the late Santonian stage and the late Campanian to the late Maastrichtian, respectively, of the Late Cretaceous. ''Kourisodon'' was originally described as a member of the "Leiodontini", more recently as a "Clidastine". Distribution ''Kourisodon puntledgensis'' is known from a single locality located in the Santonian Pender Formation along the banks of the Puntledge River (after which it was named). ''K. puntledgensis'' was a small mosasaur, with an estimated length of about 3.75 meters (around 12.3 feet). It shared its environment with a variety of Elasmosaurids, turtles, and other mosasaurs, althou ...
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Kourisodon
''Kourisodon'' ("razor tooth") is an extinct genus of mosasaur. Fossils have been found from Vancouver Island in British Columbia, Canada,Nicholls, E. L. and Meckert, D. (2002). Marine reptiles from the Nanaimo Group (Upper Cretaceous) of Vancouver Island. ''Canadian Journal of Earth Science'' 39(11):1591-1603. as well as from the Izumi Group of Japan. These finds date back to the late Santonian stage and the late Campanian to the late Maastrichtian, respectively, of the Late Cretaceous. ''Kourisodon'' was originally described as a member of the "Leiodontini", more recently as a "Clidastine". Distribution ''Kourisodon puntledgensis'' is known from a single locality located in the Santonian Pender Formation along the banks of the Puntledge River (after which it was named). ''K. puntledgensis'' was a small mosasaur, with an estimated length of about 3.75 meters (around 12.3 feet). It shared its environment with a variety of Elasmosaurids, turtles, and other mosasaurs, althou ...
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Kourisodon Puntledgensis Life Reconstruction
''Kourisodon'' ("razor tooth") is an extinct genus of mosasaur. Fossils have been found from Vancouver Island in British Columbia, Canada,Nicholls, E. L. and Meckert, D. (2002). Marine reptiles from the Nanaimo Group (Upper Cretaceous) of Vancouver Island. ''Canadian Journal of Earth Science'' 39(11):1591-1603. as well as from the Izumi Group of Japan. These finds date back to the late Santonian stage and the late Campanian to the late Maastrichtian, respectively, of the Late Cretaceous. ''Kourisodon'' was originally described as a member of the "Leiodontini", more recently as a "Clidastine". Distribution ''Kourisodon puntledgensis'' is known from a single locality located in the Santonian Pender Formation along the banks of the Puntledge River (after which it was named). ''K. puntledgensis'' was a small mosasaur, with an estimated length of about 3.75 meters (around 12.3 feet). It shared its environment with a variety of Elasmosaurids, turtles, and other mosasaurs, althou ...
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Pender Formation
The Pender Formation is a geologic formation in British Columbia. It preserves fossils dating back to the Cretaceous period. Organisms that have been recovered from this formation include indeterminate Elasmosaurids, Mosasaurids, and a Chelonioidean, as well as the sea turtle ''Desmatochelys'' and the Mosasaurine ''Kourisodon''. See also * List of fossiliferous stratigraphic units in British Columbia This is a list of fossiliferous stratigraphic units in British Columbia, Canada. References * {{Lists of fossiliferous stratigraphic units in Canada British Columbia British Columbia (commonly abbreviated as BC) is the westernmost prov ... References References * Cretaceous British Columbia {{British Columbia-stub ...
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Santonian
The Santonian is an age in the geologic timescale or a chronostratigraphic stage. It is a subdivision of the Late Cretaceous Epoch or Upper Cretaceous Series. It spans the time between 86.3 ± 0.7 mya (million years ago) and 83.6 ± 0.7 mya. The Santonian is preceded by the Coniacian and is followed by the Campanian.Gradstein ''et al.'' (2004) Stratigraphic definition The Santonian Stage was established by French geologist Henri Coquand in 1857. It is named after the city of Saintes in the region of Saintonge, where the original type locality is located. The base of the Santonian Stage is defined by the appearance of the inoceramid bivalve ''Cladoceramus undulatoplicatus''. The GSSP (official reference profile) for the base of the Santonian Stage is located near Olazagutia, Spain; it was ratified by the Subcommission on Cretaceous Stratigraphy in 2012. The Santonian's top (the base of the Campanian Stage) is informally marked by the extinction of the crinoid '' Marsupites tes ...
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Mosasaurs Of North America
Mosasaurs (from Latin ''Mosa'' meaning the 'Meuse', and Greek ' meaning 'lizard') comprise a group of extinct, large marine reptiles from the Late Cretaceous. Their first fossil remains were discovered in a limestone quarry at Maastricht on the Meuse in 1764. They belong to the order Squamata, which includes lizards and snakes. Mosasaurs probably evolved from an extinct group of aquatic lizards known as aigialosaurs in the Earliest Late Cretaceous with 42 described genera. During the last 20 million years of the Cretaceous period (Turonian–Maastrichtian ages), with the extinction of the ichthyosaurs and pliosaurs, mosasaurs became the dominant marine predators. They themselves became extinct as a result of the K-Pg event at the end of the Cretaceous period, about 66 million years ago. Description Mosasaurs breathed air, were powerful swimmers, and were well-adapted to living in the warm, shallow inland seas prevalent during the Late Cretaceous period. Mosasaurs were so we ...
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Mosasaurids
Mosasaurs (from Latin ''Mosa'' meaning the 'Meuse', and Greek ' meaning 'lizard') comprise a group of extinct, large marine reptiles from the Late Cretaceous. Their first fossil remains were discovered in a limestone quarry at Maastricht on the Meuse in 1764. They belong to the order Squamata, which includes lizards and snakes. Mosasaurs probably evolved from an extinct group of aquatic lizards known as aigialosaurs in the Earliest Late Cretaceous with 42 described genera. During the last 20 million years of the Cretaceous period (Turonian–Maastrichtian ages), with the extinction of the ichthyosaurs and pliosaurs, mosasaurs became the dominant marine predators. They themselves became extinct as a result of the K-Pg event at the end of the Cretaceous period, about 66 million years ago. Description Mosasaurs breathed air, were powerful swimmers, and were well-adapted to living in the warm, shallow inland seas prevalent during the Late Cretaceous period. Mosasaurs were so we ...
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Paleobiology Database
The Paleobiology Database is an online resource for information on the distribution and classification of fossil animals, plants, and microorganisms. History The Paleobiology Database (PBDB) originated in the NCEAS-funded Phanerozoic Marine Paleofaunal Database initiative, which operated from August 1998 through August 2000. From 2000 to 2015, PBDB received funding from the National Science Foundation. PBDB also received support form the Australian Research Council. From 2000 to 2010 it was housed at the National Center for Ecological Analysis and Synthesis, a cross-disciplinary research center within the University of California, Santa Barbara. It is currently housed at University of Wisconsin-Madison and overseen by an international committee of major data contributors. The Paleobiology Database works closely with the Neotoma Paleoecology Database, which has a similar intellectual history, but has focused on the Quaternary (with an emphasis on the late Pleistocene and Holocen ...
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Canadian Fossil Discovery Centre
The Canadian Fossil Discovery Centre, formerly known as the Morden and District Museum, is located in Morden, Manitoba in the lower level of the Access Events Centre. The museum currently houses the largest collection of marine reptile fossils in Canada. (Janzic and Hatcher 2008; Nicholls 1988) Collections Field excavations occur along the Manitoba Escarpment area of the Pembina Hills in the Red River Valley. Most fossils that are collected are from the Pembina and Millwood Members of the Pierre Shale. The Discovery Centre also houses numerous specimens of the Cretaceous squid '' Tusoteuthis longa''; these specimens represent the northernmost known occurrence of these molluscs in the Cretaceous seas of North America. (Nicholls and Isaak, 1987). The centre houses the world's largest publicly exhibited mosasaur of the species '' Tylosaurus pembinensis'', Bruce. Bruce received a Guinness World Record in on August 22, 2014 and was featured in the 2015 Guinness Book of World Re ...
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Shikoku
is the smallest of the four main islands of Japan. It is long and between wide. It has a population of 3.8 million (, 3.1%). It is south of Honshu and northeast of Kyushu. Shikoku's ancient names include ''Iyo-no-futana-shima'' (), ''Iyo-shima'' (), and ''Futana-shima'' (), and its current name refers to the four former provinces that made up the island: Awa, Tosa, Sanuki, and Iyo. Geography Shikoku Island, comprising Shikoku and its surrounding islets, covers about and consists of four prefectures: Ehime, Kagawa, Kōchi, and Tokushima. Across the Seto Inland Sea lie Wakayama, Osaka, Hyōgo, Okayama, Hiroshima, and Yamaguchi Prefectures on Honshu. To the west lie Ōita and Miyazaki Prefectures on Kyushu. Shikoku is ranked as the 50th largest island by area in the world. Additionally, it is ranked as the 23rd most populated island in the world, with a population density of 193 inhabitants per square kilometre (500/sq mi). Mountains running east and west d ...
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Pacific Ocean
The Pacific Ocean is the largest and deepest of Earth's five oceanic divisions. It extends from the Arctic Ocean in the north to the Southern Ocean (or, depending on definition, to Antarctica) in the south, and is bounded by the continents of Asia and Oceania in the west and the Americas in the east. At in area (as defined with a southern Antarctic border), this largest division of the World Ocean—and, in turn, the hydrosphere—covers about 46% of Earth's water surface and about 32% of its total surface area, larger than Earth's entire land area combined .Pacific Ocean
. '' Britannica Concise.'' 2008: Encyclopædia Britannica, Inc.
The centers of both the

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Polycotylidae
Polycotylidae is a family of plesiosaurs from the Cretaceous, a sister group to Leptocleididae. Polycotylids first appeared during the Albian stage of the Early Cretaceous, before becoming abundant and widespread during the early Late Cretaceous. Several species survived into the final stage of the Cretaceous, the Maastrichtian. With their short necks and large elongated heads, they resemble the pliosaurs, but closer phylogenetic studies indicate that they share many common features with the Leptocleididae and Elasmosauridae. They have been found worldwide, with specimens reported from New Zealand, Australia, Japan, Morocco, the US, Canada, Eastern Europe, and South America. Phylogeny Cladogram A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to d ... after Albright, Gillette and Tit ...
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Elasmosauridae
Elasmosauridae is an extinct family of plesiosaurs, often called elasmosaurs. They had the longest necks of the plesiosaurs and existed from the Hauterivian to the Maastrichtian stages of the Cretaceous, and represented one of the two groups of plesiosaurs present at the end of the Cretaceous alongside Polycotylidae. Their diet mainly consisted of crustaceans and molluscs. Description The earliest elasmosaurids were mid-sized, about . In the Late Cretaceous, elasmosaurids grew as large as , such as '' Styxosaurus'', ''Albertonectes'', and '' Thalassomedon''. Their necks were the longest of all the plesiosaurs, with anywhere between 32 and 76 (''Albertonectes'') cervical vertebrae. They weighed up to several tons. Classification Early three-family classification Though Cope had originally recognized ''Elasmosaurus'' as a plesiosaur, in an 1869 paper he placed it, with ''Cimoliasaurus'' and ''Crymocetus'', in a new order of sauropterygian reptiles. He named the group Streptos ...
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